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A new species of machaeroidine creodont, Machaeroides simpsoni, is described from the Lostcabinian and Gardnerbuttean of the Wind River Formation, Natrona County, Wyoming. The sabertooth adaptation is fully developed in this species, which has a combination both of more primitive characters and a more derived sabertooth condition than in the Bridgerian species, M. eothen. Some morphological features of Machaeroides indicate that the machaeroidines are more closely allied with the oxyaenids than with the limnocyonids.

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