While savoring your double mushroom pizza or contemplating the increasing number of mold spots on your bathroom wall, have you ever wondered how these organisms evolved and how long they have been around? Well, we have, and constantly do! Surprisingly, however, obtaining satisfying answers to these questions is difficult. Molecular-clock estimates have greatly varied as to the origin of the fungi (e.g., Heckman et al., 2001; Taylor and Berbee, 2006). Likewise the fossil record has been inconclusive with regard to the origin of the fungi and is astonishingly incomplete relative to the evolutionary history of the various fungal...

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