In their comment on our paper, Wilson and Palmer (2001) suggest that the borings in the Richmondian Onniella epibole are all domichnial and, thus, post mortem with respect to the life of the brachiopod. We originally had argued that the mixed indicators of boring habit could be explained neither by site-selective boring alone nor by site-non-selective boring alone. In addition, we found that post mortem (i.e., post-disarticulation, site-non-selective) boring could not adequately explain the evidence, especially the valve stereotypy. Instead, a mixture of boring habits seemed to be required. Whether such a postulation of multiple mechanisms constitutes an “unnecessary multiplication”...

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