The microfacies and statistical study of a Middle Devonian bioherm has revealed that, unlike many similar structures, the investigated one is a composite body in which a transgressive phase of stromatoporoidal growth overlying dolomites has been welded by a horn coral and echinoderm development to a regressive phase of stromatoporoidal growth which is itself overlain by another dolomitic zone. The ecological zones resulting from this strikingly cyclical evolution were found to be, in increasing relative depth order: matlike stromatoporoids, horn corals, and arenaceous Foraminifera, echinoderms, and ostracods.

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