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An integrated study of trace fossil assemblages from Middle Triassic deposits of the eastern border of the French Massif Central including analysis of primary sedimentary structures plant imprints invertebrate lebensspuren and vertebrate trackways has enabled reconstruction of the depositional environments. In this study, statistical methods are used to support morphological observations, permitting better definition of the ichnotaxa formed by reptilian vertebrates In addition by applying other statistical methods and the concept of “Euclidian distance” it is possible to estimate population sizes for the trackmakers. Using these data a reconstruction of the biological relationships between the different trackmakers represented by tracks in the trackway areas and between the trackmakers and their environment is proposed.

Lithology primary sedimentary structures and lebensspuren indicate that the trackway areas of this Triassic borderland were large sandy shores at the edge of a sea orlagoon The succession of footprint bearing beds in the lower sandstones or “Grés inférieurs du Lyonnais,” suggests cycles of deposition due to repeated marine transgressions.

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