Abstract The papers in this volume concentrate on the terminal Palaeozoic and Mesozoic non-marine formations of Thailand and Laos. Similar formations are also known from other countries in mainland SE Asia, such as Malaysia (where several non-marine plant-bearing formations of Jurassic to Cretaceous age are known: Lee et al. 2004), Vietnam and Cambodia (where red beds similar to those of the Khorat Group of Thailand are widespread: Workman 1977; Fontaine & Workman 1978). However, the non-marine Late Palaeozoic and Mesozoic fossils of Thailand and Laos have received much more attention than those of neighbouring countries, from which little has been reported. What we know of the Late Palaeozoic and Mesozoic ecosystems of SE Asia is therefore very largely based on the Thai and Lao records. The study of these Late Palaeozoic and Mesozoic non-marine assemblages began in the 1890s when French geologists took part in expeditions to the Lao principalities that served both a scientific and a political purpose (the latter being clearly predominant, as the more or less avowed aim was to bring that area under French influence). Counillon's 1896 report of a dicynodont skull from the vicinity of Luang Prabang thus is a landmark in the history of vertebrate palaeontology in SE Asia

Abstract Trees are an obvious component of most landscapes. Artists' views of Mesozoic landscapes regularly feature modern trees such as firs ( Abies and Picea ) and monkey-puzzle trees ( Araucaria araucana ). However, these reconstructions are highly hypothetical and, in reality, very little is known about the silhouettes of Mesozoic trees. In Mesozoic (Middle Jurassic to Early Cretaceous) strata of Thailand, large conifer logs with different types of architecture are evident, a rare opportunity for architectural studies. Various methods to estimate the original diameter and height of the trees are assessed. Among our material some trees have the classical Christmas-tree shape, whereas others are more oak-like in silhouette. All these trees lived in forest environments. Tree shape is strongly related to environment, and is still under-used as a palaeoecological tool. Reconstructing trees and vegetation has wide implications, from evaluating dinosaur herbivory to calculating elements of the carbon cycle.

Abstract We describe a new elasmobranch fauna from the lower part of the Khlong Min Formation in Thailand. The fauna includes Hybodus sp., Asteracanthus sp., Lonchidion reesunderwoodi sp. nov., Belemnobatis aominensis sp. nov., and possibly a second species of Belemnobatis. This fauna supports a Bathonian–Callovian age for the Khlong Min Formation, and suggests a close taxonomic relationship between the Middle Jurassic elasmobranch faunas of Europe and Thailand. The presence of a monolayered enameloid in Belemnobatis aominensis sp. nov. and other primitive batoids is interpreted as the retention of a primitive character for neoselachians, which would suggest a divergence time between the batoids and the rest of the neoselachian sharks as early as the Carboniferous–Permian boundary.

Abstract Most Mesozoic vertebrate species are represented by scarce and incomplete specimens, preventing statistical studies of morphometric features. Moreover, rich vertebrate assemblages are rarely excavated in conditions that allow taphonomical studies. Lepidotes buddhabutrensis is a common species found in the Late Jurassic–Early Cretaceous locality of Phu Nam Jun, Phu Kradung Formation, in NE Thailand. Individuals, collected during systematic excavation since 2002, show great variations in preservation states and body postures. In this paper we study the mode of variation of morphometric features of the fish population, the growth mode, and the relationship between morphology and size. We assess the range of variation in preservation and taphonomy, based on arbitrarily defined scales, to test if vertical variations occur in the sample of individuals within the site. We test possible favoured orientation of specimens within the assemblage. In contrast to preliminary field observations, statistical analyses show that all individuals belong to a single Gaussian population and that gross morphological shape variations are related only to size during fish growth. L. buddhabutrensis shows a positive allometric growth for the pectoral to dorsal, and pectoral to anal fin distances, and a negative allometric growth for the unpaired fins (dorsal and anal fins lengths). We detected no relationships between the vertical location of the fishes within the fossiliferous deposit and the body shape of the specimens, nor between the state of preservation and the taphonomy, but there are significant differences in the state of preservation according to the position of the fishes in the fossiliferous deposit. The occurrence of a single Gaussian population and the absence of morphological and preservational variations through the depositional column are evidence that the fish assemblage is probably the result of a single mass mortality event. The apparent diversity in morphology is probably due to variations in the mode of preservation. The fish appear to have been oriented by a current at the time of deposition at the top of the fossiliferous deposit only.

Abstract This first overview of the bony fish record from the Jurassic and Cretaceous continental deposits of Thailand reveals a significant diversity, with 16 taxa in four formations (the Khlong Min, Phu Kradung, Sao Khua and Khok Kruat Fms). Four of these taxa have already been diagnosed and described, and a couple of others are sufficiently well preserved to be diagnosed in the future. The other taxa are represented at present by fragmentary and isolated remains. The highest diversity is observed among ‘semionotids’, which occur in the four formations. Sinamiids are represented by at least three taxa that occur only in the Sao Khua and the Khok Kruat Formations. Pycnodont fishes are known by rare and isolated dentitions and teeth in the Khlong Min and Sao Khua Formations, and lungfishes referred to Ferganoceratodus occur in the Khlong Min and the Phu Kradung Formations. The assemblages provide few palaeogeographical indications at present, except for evidence of relationships with China and Central Asia. However, it is expected than once the phylogenetic relationships of these taxa are resolved, we will be able to reconstruct precise palaeogeographical scenarios.

Abstract The turtle assemblages from the Khorat Group consist mainly of trionychoids. They include the primitive Trionychoidae Basilochelys and basal eucryptodiran turtles from the Phu Kradung Formation (?Late Jurassic); the adocid Isanemys srisuki , the carettochelyid Kizylkumemys sp. and undetermined Trionychoidea from the Sao Khua Formation (Early Cretaceous); and the carettochelyid Kizylkumemys khoratensis and the adocid Shachemys sp. from the Khok Kruat Formation (Aptian). Our study shows some faunal links between the turtle faunas from the Khorat Group and those from the peripheral regions of Asia during the time span of the Khorat Group. Thus the coastal regions of Asia, and more particularly SE Asia, may have been important places for the origin and early diversification of the trionychoids.

Abstract A large cryptodiran turtle, Basilochelys macrobios n. gen. n. sp. is described from the latest Jurassic–earliest Cretaceous Phu Kradung Formation of NE Thailand, on the basis of skull, shell and other postcranial elements. Basilochelys presents a combination of primitive and derived characters. The derived characters include sculptured skull roof and shell surface; deeply embedded canalis caroticus internus; foramen posterius canalis carotici interni completely surrounded by pterygoid; neural formula of 6>4<6<6<6<6; anteroposteriorly expanded eleventh and twelfth marginal scutes extending onto the suprapygal and costal plates; narrow vertebral scutes; plastron sutured to the carapace, with large and wide anterior and posterior lobes, long and narrow bridge, very narrow axillary and inguinal notch; wide entoplastron; humeropectoral sulcus located on the posterior part of the entoplastron; anal notch absent. This taxon is placed in Trionychoidae and considered as the most basal member of that group.

Abstract Bone histology is the most comprehensive way of obtaining data on growth and life history for dinosaurs. Humeri and femora of the basal titanosaur Phuwiangosaurus sirindhornae from the Early Cretaceous of Thailand were sampled by core drilling. The sample represents growth series with humeri ranging in size from 71.0 to 110.0 cm and femora ranging in size from 38.5 to 112.0 cm. The bone tissue is continuously growing laminar fibro-lamellar bone typical for virtually all sauropods. Several ontogenetic stages can be distinguished, and a general growth pattern is deduced on the basis of different-sized individuals. Humeri differ from femora in generally showing more remodelling by secondary osteons.

Abstract Postcranial remains of a small theropod dinosaur, including vertebrae, incomplete pubes, tibiae, an incomplete fibula, metatarsals and phalanges, from the Early Cretaceous Sao Khua Formation of Phu Wiang, Khon Kaen Province, NE Thailand, are described as a new taxon of ornithomimosaur, Kinnareemimus khonkaenensis , gen. et sp. nov. This early ‘ostrich dinosaur’ is characterized by a fairly advanced metatarsus, in which metatarsal III, although still visible proximally between metatarsals II and IV in cranial view, is markedly ‘pinched’ more distally and becomes triangular in cross-section. The condition of its metatarsus shows that Kinnareemimus khonkaenensis is more derived than the geologically younger primitive ornithomimosaurs Harpymimus and Garudimimus , but less derived than Archaeornithomimus . Its occurrence in the Early Cretaceous of Thailand suggests that advanced ornithomimosaurs may have originated in Asia.

Abstract Vertebrate footprints have been discovered in recent years from seven Mesozoic formations of Thailand and Laos dating from the Late Triassic (Kuchinari Group) and the Early Cretaceous (Khorat Group). The sites are reviewed here in chronological order. The ichnological record reflects fairly well the broad picture of the evolution of continental vertebrates in Asia known from the skeletal record. Norian basal archosaurs are replaced by Rhaetian dinosaurs although both footprint morphotypes look different from the contemporaneous European and North American forms. Two successive ornithopod radiations can be observed in the Early Cretaceous, with primitive small tetradactyl Hypsilophodon -like dinosaurs in the Earliest Cretaceous followed by advanced iguanodontoids with tridactyl fleshy footprints in the Aptian. Late Early Cretaceous dinosaur footprints from NE Thailand, however, do not validate previous hypotheses on the geographical distribution of Cretaceous ornithopod tracks in Asia. The ichnological record also reveals a hitherto unsuspected high diversity of theropods in the early Cretaceous with many different morphotypes.

Abstract Phosphatic remains (tooth enamel, turtle shell fragments and fish scales) of continental vertebrates (freshwater fish, crocodilians, turtles, and theropod and sauropod dinosaurs) recovered from eight localities of NE Thailand ranging in age from the Late Jurassic to the late Early Cretaceous have been analysed for their oxygen isotopic compositions (δ 18 O p ). From these preliminary data, local meteoric water δ 18 O w values estimated using δ 18 O p values of crocodilians and turtles range from −4.1±2‰ at the end of the Jurassic to −8.3±2‰ during the Early Cretaceous, suggesting a transition from dry to wetter climates with increasing amount of seasonal precipitation from several hundred millimetres per year to several thousand millimetres. Measurable offsets in δ 18 O p values observed between dinosaur taxa (the spinosaurid theropod Siamosaurus , other theropods and nemegtosaurid sauropods) are interpreted in terms of differences in water strategies, and suggest that Siamosaurus had habits similar to those of semi-aquatic vertebrates such as crocodilians or freshwater turtles.

Abstract Over the last 25 years, rich vertebrate assemblages have been discovered in three distinct formations of the Late Jurassic and Early Cretaceous of Thailand. This work aims to compare the taxonomic assemblages of SE Asia within their palaeogeographical context in Asia. Occurrences of 477 taxa in 94 Regional Faunal Assemblages (RFA) have provided the raw material for producing a dissimilarity matrix based on the Raup & Crick index. These distances have been investigated statistically to infer relationships between the diverse faunal assemblages in space and time. Our results show that the Thai formations are more similar to each other than to any other formations, suggesting a strong provincialism. The relationship of SE Asian RFAs with other Asian RFAs is more influenced by the presence of freshwater or near-shore taxa than by strictly terrestrial ones. Our analysis shows that the faunal interchange between RFAs was rather low from the Late Jurassic to the end of the Early Cretaceous. However, faunal dispersals dramatically decreased during the mid-Early Cretaceous in Asia. The faunas show an overall stronger provincialism during the mid-Early Cretaceous, indicating the role of possible geographical barriers. This event is characterized by the absence of ornithischian dinosaurs in the Sao Khua Formation although they are present in the under- and overlying formations. Taxonomic diversity and exchanges between faunal assemblages recovered rapidly as early as the Aptian in Asia, but the fauna of SE Asia still retained a strong biogeographical signature.