Abstract In Scotland and Ireland, a Laurentian passive margin sequence, the Dalradian Supergroup, was deformed during the c. 470–460 Ma Grampian orogeny, resulting in the formation of crustal-scale recumbent nappes. In Ireland, this passive margin sequence is in general bounded to the SE by the Fair Head–Clew Bay Line (FHCBL), a segment of a major lineament within the Caledonides. Adjacent to the FHCBL, Dalradian metasediments in two separate inliers have undergone post-Grampian strike–slip movement, with the initially flat-lying Grampian nappe fabric acting as a décollement-like slip surface in both cases. As the orientation of these foliation slip surfaces was oblique to the local shear plane in both inliers, displacement along these pre-existing foliation surfaces was also accompanied by crenulation slip. However, the crenulation-slip morphologies produced imply the opposite sense of movement in the two inliers. 40 Ar− 39 Ar dating of muscovite defining the crenulation-slip surfaces indicates that post-Grampian dextral displacement took place along the FHCBL at 448 ± 3 Ma. A subsequent phase of sinistral movement along the FHCBL took place at c. 400 Ma, based on previously published Rb–Sr muscovite ages for synkinematic pegmatites. The kinematic information obtained from crenulationslip morphologies combined with geochronology can thus be used to constrain the reactivation history of a major crustal-scale shear zone.