Abstract In Europe, North Africa and Asia Minor, the remains of Pennsylvanian sedimentary basins bearing continental deposits either intimately mixed with shallow-marine strata or deposited in exclusively continental settings are preserved. Long-lasting research on these basins allowed the definition of regional stages and substages based on marine fauna and terrestrial flora, later extended by terrestrial and freshwater faunal biostratigraphies. Glacioeustatically driven marine bands provide laterally widespread correlation markers; however, where such bands are missing only biostratigraphic control exists. Resolution of biostratigraphic zonations combined with gaps in sedimentary successions and variable quality of the fossil record throughout the basin fills do not allow in all cases a precise correlation between the Pennsylvanian basins in Europe and, in turn, the timing of tectonic, climatic and biotic events, and thus an absolute complete understanding of the response of terrestrial and freshwater biota to climate changes across eastern tropical Pangaea. A helpful tool is new radioisotopic ages of intercalated volcaniclastics that reveal the partial diachroneity of some widely used biostratigraphies. We attempt to present the current state of the art to stimulate further research to mitigate gaps in our knowledge.

Abstract We present an updated look at Carboniferous brachiopod biozonation from most of the world framed into a revised Carboniferous palaeogeography, based on a selection of the literature published on Carboniferous brachiopods since the nineteenth century. The biostratigraphic significance of the most important brachiopod taxa is synthesized in seven geographical correlations. The Mississippian is characterized by rich brachiopod faunas, with widespread taxa with a good potential for global correlation, such as Rugosochonetes , Delepinea , Buxtonia , Antiquatonia , Spinocarinifera , Marginatia , Fluctuaria , Ovatia , Rhipidomella , Lamellosathyris , Unispirifer , Tylothyris and Syringothyris . From the mid-Visean to the late Serpukhovian, taxa of gigantoproductidines are biostratigraphically significant, and occur everywhere except South America and Australia, which remain as distinct faunal successions for most of the period. A major turnover occurs at the beginning of the Pennsylvanian, characterized by a higher degree of provincialism. Pennsylvanian brachiopod faunas are diverse in China, Russia and North America, but otherwise they are less developed and are characterized mostly by endemic taxa, hampering long-distance correlation. An exception is the rapid diversification of taxa of the Choristitinae, which were widespread from the Bashkirian to the Moscovian, allowing long-distance correlation.

Abstract During the Carboniferous, crinoids were commonly so abundant that their skeletal ossicles formed limestones termed encrinites. Major evolutionary changes occurred within the Camerata and Articuliformes, as the former were displaced by the latter as the dominant clade. Both the Mississippian and the Pennsylvanian subperiods started with high evolutionary rates and ended with low evolutionary rates associated with glaciation. Although not typically used for biostratigraphy, crown-based crinoid genera can be used as biostratigraphic indicators for Carboniferous stages. Paleozoic crinoid biodiversity reached its maximum during the Carboniferous, from which there are numerous well-documented localities with high biodiversity. Faunas from the palaeobiogeographical regions of Laurussia, Palaeo-Tethys and Gondwana are reviewed. For Mississippian crinoids, 37 genera are designated as biostratigraphically useful; and, for the Pennsylvanian, 44 genera are identified. Recognition of the utility of these genera for biostratigraphy is important for dating crinoid deposits, which may be devoid of other biostratigraphically useful fossils, and add to our overall ability to delineate the temporal resolution of life on Earth.

Abstract Rugose corals are one of the major fossil groups in shallow-water environments. They played an important role in dividing and correlating Carboniferous strata during the last century, when regional biostratigraphic schemes were established, and may be useful for long-distance correlation. Carboniferous rugose corals document two evolutionary events. One is the Tournaisian recovery event, with abundant occurrences of typical Carboniferous rugose corals such as columellate taxa and a significant diversification of large, dissepimented corals. The other is the changeover of rugose coral composition at the mid-Carboniferous boundary, which is represented by the disappearance of many large dissepimented taxa with complex axial structures and the appearance of typical Pennsylvanian taxa characterized by compound rugose taxa. The biostratigraphic scales for rugose corals show a finer temporal resolution in the Mississippian than in the Pennsylvanian, which was probably caused by the Late Paleozoic Ice Age that resulted in glacial–eustatic changes and a lack of continuous Pennsylvanian carbonate strata. The Pennsylvanian rugose corals are totally missing in the Cimmerian Continent. High-resolution biostratigraphy of rugose corals has so far only been achieved in few regions for the Mississippian timescale. In most regions, more detailed taxonomic work and precise correlations between different fossil groups are needed.

Abstract Considerable progress has been made by international teams in refining the traditional ammonoid zonation that remains the backbone of Carboniferous stratigraphy. The Carboniferous ammonoid genozones, with a few gaps, are now recognized throughout the entire system in most successions worldwide. Refined collecting and documentation of occurrences in Western Europe, North Africa, the Urals, China and North America aimed to establish the first evolutionary occurrences, and facilitated correlation with foraminiferal and conodont scales for most of the Carboniferous. From ten to eleven ammonoid genozones are now recognized in the Mississippian, and eight to nine genozones in the Pennsylvanian. Of these, the established lower boundaries of the subsystems are reasonably well correlated with the ammonoid zonation, whereas correlations with the ratified foraminiferal-based lower boundary of the Visean and other stage boundaries, currently under discussion, need further research. Future success in the ammonoid geochronology will also depend on accurate identification and re-illustration of the type material, including material described by pioneers of ammonoid biostratigraphy.

Abstract Carboniferous conodont biostratigraphy comprises regional zonations that reflect the palaeogeographical distribution of taxa and distinct shallow-water and deep-water conodont biofacies. Some species have a global distribution and can effect high quality correlations. These taxa are incorporated into definitions of global Carboniferous chronostratigraphic units. A standard global Carboniferous zonation has not been developed. The lowermost Mississippian is zoned by Siphonodella species, excepet in shallow-water facies, where other polygnathids are used. Gnathodus species radiated during the Tournaisian and are used to define many Mississippian zones. A late Tournaisian maximum in diversity, characterized by short-lived genera, was followed by lower diversity faunas of Gnathodus species and carminate genera through the Visean and Serpukhovian. By the late Visean and Serpukhovian, Lochriea provides better biostratigraphic resolution. Shallow-water zonations based on Cavusgnathus and Mestognathus are difficult to correlate. An extinction event near the base of the Pennsylvanian was followed by the appearance of new gnathodid genera: Rhachistognathus , Declinognathodus , Neognathodus , Idiognathoides and Idiognathodus . By the middle of the Moscovian, few genera remained: Idiognathodus , Neognathodus and Swadelina. During the middle Kasimovian and Gzhelian, only Idiognathodus and Streptognathodus species were common. Near the end of the Gzhelian, a rediversification of Streptognathodus species extended into the Cisuralian.

Abstract Among several groups of fishes existing in the Carboniferous, the Chondrichthyes appear to have the greatest stratigraphic potential. However, despite the long history of investigation into Paleozoic sharks, and especially their teeth, our knowledge of their usefulness in biostratigraphy and palaeoecology is still at an early stage of development. This is mainly because for a long time palaeoichthyologists have been focused on descriptions of individual taxa, and not on documenting whole assemblages. The microscopic teeth of pelagic stem-group Chondrichthyes, such as Thrinacodus (Phoebodontiformes), Denaea and Stethacanthulus (Falcatidae, Symmoriiformes) appear to be more useful than macrofossils (e.g. tooth plates of Holocephali) because of their wider geographical distribution and weaker facies dependence.

Abstract This paper presents a summary of palynological data for Pennsylvanian age coal beds in the Appalachian Basin, discussed primarily from a biostratigraphic perspective. Coal bed palynofloras of Lower Pennsylvanian through early Permian age are compared and correlated with miospore assemblage zones established for western Europe, and the Eastern Interior (Illinois) and Western Interior Basins of the mid-continent USA. Lower Pennsylvanian palynofloras, which are dominated by lycopsid spores, are correlative with the Langsettian of western Europe and the Morrowan of the Eastern and Western Interior mid-continent USA Basins. Stratigraphically useful palynotaxa include Dictyotriletes bireticulatus , Radiizonates striatus , Schulzospora rara , Granasporites medius , Laevigatosporites minor and Endosporites globiformis . Middle Pennsylvanian palynofloras change through time, being lycopsid dominant in the lower part and more heterogeneous in the middle and upper parts with increased contributions from other Pennsylvanian plant groups. They are correlative with the Duckmantian, Bolsovian and Asturian of western Europe and the Atokan and Desmoinesian of the Eastern and Western Interior mid-continent USA Basins. Stratigraphically useful palynotaxa include Secarisporites remotus , Microreticulatisporites sulcatus , Vestispora fenestrata , Triquitrites sculptilis , Laevigatosporites globosus , Radiizonates difformis , Torispora securis , Triquitrites minutus , Mooreisporites inusitatus , Murospora kosankei , Thymospora pseudothiessenii and Schopfites dimorphus . Upper Pennsylvanian and lower Permian coal beds in the Appalachian Basin, in contrast to their Lower and Middle Pennsylvanian counterparts, are strongly dominated by tree fern spore palynotaxa. Palynofloras correlate with the Stephanian and Autunian of western Europe and the Missourian, Virgilian and Wolcampian of the Eastern and Western Interior mid-continent USA Basins.

Abstract In the Carboniferous, terrestrial vegetation became widespread, diverse and abundant. The resulting fossil record has proved to be an effective biostratigraphic tool for intra- and interbasinal correlations. Besides palaeogeographical configurations, Carboniferous plant biostratigraphy is affected by a transition from greenhouse conditions during most of the Mississippian to an icehouse climate in the Pennsylvanian. The greenhouse Mississippian climate resulted in weak provincialism, with a cosmopolitan flora ranging from the tropics to middle latitudes. The global cooling around the Mississippian–Pennsylvanian boundary enhanced development of a latitudinal climatic zonation and related floral provincialism. These changes are expressed in the recognition of distinct realms or kingdoms, where the tropical Amerosinian Realm (or Euramerican and Cathaysian realms) is surrounded by the Angaran and Gondwanan realms occupying middle to high latitudes of the northern and southern hemispheres, respectively. Floristic endemism in the Pennsylvanian precludes development of a global macrofloral biostratigraphy. Instead, each realm or area has its own biostratigraphic scheme. Poorer and less diverse floras of the Gondwanan and Angaran realms resulted in the establishment of relatively low-resolution macrofloral biostratigraphic schemes. Higher-resolution macrofloral zonations exist only in the tropical Amerosinian Realm due to diverse and abundant floras dominated by free-sporing and early seed plants occupying extensive wetlands.

Abstract For the biostratigraphy of mixed continental-marine and purely continental sections in the palaeotropical belt of Euramerica, 9 insect and 8 conchostracan zones are newly defined or improved. These zones encompass the time interval from the Early Pennsylvanian (middle Bashkirian) up into the early Permian (early Asselian) of the Euramercian biotic province. They are linked as much as possible to the marine Standard Global Chronostrigraphic Scale by common occurrences of insects and/or conchostracans with conodonts in mixed marine-continental sections as well as by the thus far available and reliable radioisotopic ages of associated volcanic rocks. This insect and conchostracan zonation is an alternative tool to the well-established macro-plant biostratigraphy of the Pennsylvanian. In contrast to the latter, only single specimens of insects or conchostracans, even if more rare than plant remains, allow biostratigraphic dating with a similar high temporal resolution.

Abstract This review summarizes research on the biostratigraphic application of Carboniferous marine and non-marine bivalves worldwide, with a focus on the southern margin of Laurussia and the Palaeotethys. Preliminary and established biostratigraphic zonations based on marine and non-marine bivalves are compiled and, if possible, correlated. Bivalve taxa of potential biostratigraphic significance are discussed, and possible limitations of bivalve zonations are outlined. Although marine and non-marine bivalves have not received primary attention for biostratigraphic purposes, a wealth of as yet unused data exists practically worldwide that can assist and complement the more favoured brachiopod, foraminiferid, ammonoid and conodont biozonations.

Abstract The Carboniferous record of tetrapod footprints is mostly of Euramerican origin and provides the basis for a footprint biostratigraphy and biochronology of Carboniferous time that identifies four tetrapod footprint biochrons: (1) stem-tetrapod biochron of Middle Devonian–early Tournaisian age; (2) Hylopus biochron of middle Tournaisian–early Bashkirian age; (3) Notalacerta–Dromopus interval biochron of early Bashkirian–Kasimovian age; and (4) Dromopus biochron of Kasimovian–early Permian age. Particularly significant is the Carboniferous tetrapod footprint record of the Maritimes basin of eastern Canada (New Brunswick, Nova Scotia and Prince Edward Island), which encompasses well-dated and stratigraphically superposed footprint assemblages of Early Mississippian–early Permian age. The Carboniferous tetrapod footprint record provides these important biostratigraphic datums: (1) oldest temnospondyls (middle Tournaisian); (2) oldest reptiliomorphs, likely anthracosaurs (middle Tournaisian); (3) oldest amniotes (early Bashkirian); and (4) oldest high-fibre herbivores (Bashkirian). Carboniferous tetrapod footprints thus provide significant insight into some major events of the Carboniferous evolution of tetrapods.

Abstract Tetrapod (amphibian and amniote) fossils of Carboniferous age are known almost exclusively from the southern part of a palaeoequatorial Euramerican province. The stratigraphic distribution of Carboniferous tetrapod fossils is used to identify five land-vertebrate faunachrons: (1) Hortonbluffian (Givetian–early Visean), the time between the first appearance datum (FAD) of tetrapods to the beginning of the Doran; (2) Doran (late Visean–early Bashkirian), the time between the FAD of the baphetid Loxomma and the beginning of the Nyranyan; (3) Nyranyan (late Bashkirian–Moscovian), the time between the FAD of the eureptile Hylonomus and the beginning of the Cobrean; (4) Cobrean (Kasimovian–late Gzhelian), the time between the FAD of the eupelycosaur Ianthasaurus and the beginning of the Coyotean; and (5) Coyotean (late Gzhelian–early Permian), the time between the FAD of the eupelycosaur Sphenacodon and the beginning of the Seymouran. This biochronology provides insight into some important evolutionary events in Carboniferous tetrapod evolution.

The Carboniferous was the time of the assembly of Pangaea by the collision of the Gondwanan and Larussian supercontinents, and the principal interval of the late Paleozoic ice ages. These tectonic and climatic events caused dramatic sea-level fluctuations and climate changes and produced a Carboniferous world that was diverse topographically and climatologically, perhaps only rivalled in that diversity by the late Cenozoic world. Furthermore, the Carboniferous was a time of the accumulation of vast coal deposits of great economic and societal significance. The temporal ordering of geological and biotic events during Carboniferous time thus is critical to the interpretation of some unique and pivotal events in Earth history. This temporal ordering is based on the Carboniferous timescale, which has been developed and refined for nearly two centuries. This book reviews the history of the development of the Carboniferous chronostratigraphic scale and includes comprehensive analyses of Carboniferous radioisotopic ages, magnetostratigraphy, isotope-based correlations, cyclostratigraphy and timescale-relevant marine and non-marine biostratigraphy and biochronology.

Abstract The Carboniferous chronostratigraphic scale consists of two subsystems, six series and seven stages. Precise numerical age control within the Carboniferous is uneven, and a global magnetic polarity timescale for the Carboniferous is far from established. Isotope stratigraphy based on Sr, C and O isotopes is at an early stage but has already identified a few Sr and C isotope events of use to global correlation. Cyclostratigraphy has created a workable astrochronology for part of Pennsylvanian time that needs better calibration. Chronostratigraphic definitions of most of the seven Carboniferous stages remain unfinished. Future research on the Carboniferous timescale should focus on Global Stratotype Section and Point (GSSP) selection for the remaining, undefined stage bases, definition and characterization of substages, and further development and integration of the Carboniferous chronostratigraphic scale with radioisotopic, magnetostratigraphic, chemostratigraphic and cyclostratigraphic tools for calibration and correlation, and the cross-correlation of non-marine and marine chronologies.

Abstract The Carboniferous chronostratigraphic scale is a hierarchy of two subsystems, six series and seven stages developed during nearly two centuries of research. Carboniferous stage nomenclature developed with the proposal of numerous regional stages/substages based primarily on palaeobotanical, foraminiferal and ammonoid biostratigraphy, especially in Western Europe, the former Soviet Union, China and the USA. From the regional stages, seven ‘global stages’ have been identified (in ascending order): Tournaisian, Visean, Serpukhovian, Bashkirian, Moscovian, Kasimovian and Gzhelian. Three of the four ratified Carboniferous GSSPs use conodont evolutionary events as the primary signal for correlation – bases of Tournaisian, Bashkirian and base of Asselian. The GSSP of the Visean base has a foraminiferal event as its primary signal. Issues in the development of a Carboniferous chronostratigraphic scale include the rank of chronostratigraphic units, provinciality, conodont biostratigraphy, palaeobotanical biostratigraphy and the development of astrochronology and other methods of chronology and correlation.