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Abstract The Ordovician was a key period in the biological and geological history of the Earth. ‘A Global Synthesis of the Ordovician System’ is presented in two volumes of The Geological Society, Special Publications series. The first volume (SP532) covers general aspects of the Ordovician and also includes the syntheses of the Ordovician successions of Europe. To provide a comprehensive global overview, this second volume (SP533) represents a journey through the Ordovician System around the world. Reviews of the Ordovician of North America include syntheses of Alaska, Greenland, Canada, the USA and Mexico, whereas the South American Ordovician is summarized in a specific chapter related to Argentina and neighbouring countries. The Ordovician System of Africa is presented in chapters covering the north and the south of the continent where significant Ordovician successions occur. Australia and New Zealand, as well as Antarctica, are visited in separate chapters. Asia provides the most complex Ordovician successions that are reviewed in chapters covering Turkey and the Levant region, the Middle East, Central Asia, Kazakhstan, India, SE Asia, China, Korea, and Japan. Our journey covers a great number of locations but, with many successions still to be fully described, our knowledge of the Ordovician of the world remains incomplete.
Abstract The Ordovician was a key period in the biological and geological history of the planet. ‘A Global Synthesis of the Ordovician System’ is presented in two volumes of The Geological Society, Special Publications . This first volume (SP532) charts the history of the Ordovician System and explores significant advances in our understanding of the period's biostratigraphy, including more precise calibration of its timescale with tephra chronology and regional alignments using astrochronology and cyclostratigraphy. Changes in the world's oceans, their shifting currents and sea levels, the biogeography of their biotas and the ambient climate are described and discussed against a background of changing palaeogeography. This first volume also includes syntheses of Ordovician geology of most European countries, including historical key areas, such as the British Isles, Baltoscandia and Bohemia.
Abstract The Ordovician successions of France and neighbouring areas of Belgium and Germany are reviewed and correlated based on international chronostratigraphic and regional biostratigraphic charts. The same three megasequences related to the rift, drift and docking of Avalonia with Baltica can be tracked in Belgium and neighbouring areas (Brabant Massif and Ardenne inliers), western (Rhenish Massif) and northeastern Germany (Rügen). The remaining investigated areas were part of Gondwana in the Ordovician. The Armorican Massif shares with the Iberian Peninsula a Furongian–Early Ordovician gap (Toledanian or Norman gap), and a continuous Mid–Late Ordovician shelf sedimentation. The Occitan Domain (Montagne Noire and Mouthoumet massifs), eastern Pyrenees and northwestern Corsica share with southwestern Sardinia continuous shelf sedimentation in the Early Ordovician, and a Mid Ordovician ‘Sardic gap’. In the Ordovician, the Maures Massif probably belonged to the same Sardo-Occitan domain. The Vosges and Schwarzwald massifs display comparable, poorly preserved Ordovician successions, suggesting affinities with the Teplá-Barrandian and/or Moldanubian zones of Central Europe.
An introduction to the Great Ordovician Biodiversification Event: insights from the Tafilalt Biota, Morocco
Abstract The exquisitely preserved, diverse and abundant fossil assemblages yielded by the ‘echinoderm meadows’ of the Tafilalt region of the eastern Anti-Atlas represent a new Konservat-Lagerstätte, one of the few exceptionally preserved Late Ordovician open-marine faunas found globally, giving us an insight into the radiation of life during the later phases of the Great Ordovician Biodiversification Event (GOBE) in high-latitude peri-Gondwana. The GOBE resulted in an unprecedented increase in the diversity of families, classes and orders, at the fastest rate of the entire Phanerozoic and represents one of the most significant events in the evolution of the marine biosphere, preceding the end-Ordovician mass extinction. Nine different phyla and several soft-bodied problematica are represented in the Tafilalt, including several notable echinoderm Lagerstätten. This volume is the culmination of over 20 years of research by several international teams and integrates a series of contributions that look at diverse aspects of the biota, including the stratigraphic distribution of the faunas, depositional environments, systematic palaeontology, preservation, palaeobiogeography and the nature and impact of the international fossil trade on these exceptionally preserved fossil faunas.
The Late Ordovician Tafilalt Biota, Anti-Atlas, Morocco: a high-latitude perspective on the GOBE
Abstract The extensive, predominantly siliciclastic deposits of the Upper Ordovician of the Tafilalt have long been the subject of scientific investigation. In the past 25 years, intensified collecting for commercial purposes has resulted in the discovery of several exceptionally-preserved faunas (Konservat-Lagerstätten) in the Tafilalt region, preserving a range of non-biomineralized and soft-bodied organisms. The preservation of these fossils in the coarse clastic sediments of the Tafilalt is surprising, and in the case of soft-bodied organisms, remarkably similar to the preservational mode of typical Ediacaran biotas. These relatively recent discoveries have increased the scientific significance of the Tafilalt Biota, providing an unparalleled insight into the composition and temporal evolution of the shallow, open-marine ecosystems and their denizens during the later stages of the Great Ordovician Biodiversification Event. At least nine different phyla, in addition to several soft-bodied problematica are represented in the Tafilalt. While the highly diverse and remarkably well-preserved echinoderm and euarthropod faunas are most emblematic for the Tafilalt Biota, further studies have revealed a relatively high diversity of molluscs and brachiopods. Among soft-bodied fossils, the problematic paropsonemid eldonids are iconic for the Tafilalt and stand out both through their abundance, and their wide temporal and geographical range throughout the area.
Ordovician stratigraphy and benthic community replacements in the eastern Anti-Atlas, Morocco
Abstract The Anti-Atlas contains a thick, volcanic-free Ordovician succession that originally deposited in a passive-margin basin. Three main sedimentary packages are bounded by major unconformities: (i) the Tremadocian–Floian Lower Fezouata and Upper Fezouata formations, which unconformably overlie a palaeorelief of Cambrian rifting volcanosedimentary complexes, and are subsequently topped by a Dapingian paraconformable gap; (ii) the Darriwilian–Katian Tachilla Formation and First Bani and Ktaoua groups, the latter unconformably overlain by a Hirnantian glaciogenic succession; and (iii) the Second Bani Group, which subsequently infilled the former glaciogenic palaeorelief. Due to the scarcity of carbonate interbeds for etching analyses, leading to rare references of conodonts, the global Ordovician chart is interpolated on the basis of microphytoplancton (acritarchs and chitinozoans), regional graptolites and brachiopods. The Ordovician counter-clockwise rotation of Gondwana led its Moroccan margin from mid- to high-latitude positions, leading to the onset of a siliciclastic, wave- and storm-dominated platform. Flooding surfaces are marked by shelly silty carbonate interbeds that reflect the episodic development of echinoderm–bryozoan meadows during Katian times; in areas protected from siliciclastic input, they reached massive and bedded bioaccumulations (Khabt-el-Hajar Formation). The Hirnantian glaciation controlled the incision of numerous tunnel channels, infilled with both alluvial to fluvial sediments and glaciomarine diamictites. The Hirnantian palaeorelief was definitively sealed during Silurian times.
Abstract Anomalocystitid mitrates represent one of the most diverse and long ranging clade of stylophorans (Early Ordovician–Middle Devonian). Although they probably originated from a peri-Gondwanan stock of early mitrocystitids during the Floian, the fossil record of anomalocystitids in the Middle–Upper Ordovician of the Mediterranean Province remains extremely scarce and largely underestimated. The unusually shaped anomalocystitid genus Diamphidiocystis was originally described in the latest Katian–Hirnantian of North America (Illinois). However, earlier occurrences of this genus in the late Darriwilian of western France (Brittany) suggest a probable peri-Gondwanan origin. Based on new Middle to Late Ordovician material from the Anti-Atlas (Morocco), Bohemia (Czech Republic) and Brittany (France), all occurrences of Mediterranean Diamphidiocystis are considered as conspecific and formally described as D. regnaulti sp. nov. The palaeobiogeographical significance of Ordovician anomalocystitid mitrates is discussed.
New Middle and Late Ordovician cornute stylophorans (Echinodermata) from Morocco and other peri-Gondwanan areas
Abstract Cornute stylophorans are a minor, although typical, component of Middle–Late Ordovician echinoderm assemblages adapted to soft siliciclastic substrates in high-latitude peri-Gondwanan regions. All previously reported occurrences of Darriwilian–Katian cornutes from the Czech Republic, France, Morocco and Spain are revised and their plate homologies reassessed. The genera Beryllia and Juliaecarpus are reinterpreted as junior synonyms of Domfrontia , and Thoralicystis is synonymized with Bohemiaecystis . Several Mediterranean scotiaecystids previously assigned to Bohemiaecystis and/or Scotiaecystis are placed within Thoralicarpus gen. nov., and cornute taxa originally left in open nomenclature by Chauvel are formally described as Bohemiaecystis chouberti sp. nov. (AVI) and Destombesicarpus izegguirenensis gen. et sp. nov. (AVIII). Other new Mediterranean taxa include Arauricystis clariondi sp. nov., Destombesicarpus budili gen. et sp. nov., Milonicystis reboulorum sp. nov., Thoralicarpus bounemrouensis gen. et sp. nov., and T . prokopi gen. et sp. nov. The six cornute genera identified in Darriwilian–Katian Moroccan echinoderm Lagerstätten are also present in coeval assemblages of at least one other Mediterranean region, thus supporting the existence of strong faunal affinities between the Anti-Atlas, the Armorican Massif, the Barrandian area and the Iberian Peninsula.
Abstract In the western Tafilalt area, eastern Anti-Atlas (Morocco), solutan echinoderms are a major faunal element of most echinoderm Lagerstätten occurring within the Lower Ktaoua Formation (late Sandbian–early Katian). For the first time, members of the class Soluta are formally described from Morocco and Africa. All solutans from the Lower Ktaoua Formation are identified as Dendrocystites aff. sedgwicki . Three size-related morphotypes, probably corresponding to successive growth stages (‘juvenile’, ‘adult’ and ‘gerontic’), could be distinguished within the abundant and well-preserved material from Morocco. The occurrence of the genus Dendrocystites in the western Tafilalt confirms the strong faunal affinities between Morocco and other regions of the Mediterranean Province (Czech Republic and Spain) in Late Ordovician times. In high palaeolatitude (peri-)Gondwanan areas, Dendrocystites was a gregarious solutan living in shallow, siliciclastic settings at or above the storm-wave base.