Hydrothermal fluids obtain organic compounds through diverse pathways. In submarine systems organic compounds are already dissolved in seawater that is heated and transformed into hydrothermal fluids through water-rock reactions. Microbes inhabiting hydrothermal systems produce metabolites that enter the fluids, and cells can be carried into the reaction zones by circulating fluids and pyrolyzed. Analogous sources of organic compounds can be anticipated in continental systems with the possible addition of novel plant- and soil-derived organic compounds from the surface. In addition, hydrothermal systems possess large potentials for abiotic organic synthesis that may add a novel suite of compounds. When sedimentary rocks...

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