The stable isotope compositions of biogenic materials record a combination of environmental parameters and biological processes. In general, the environment provides a range of isotopic compositional inputs, and an animal processes those signals through dietary preference, physiology, behavior, etc. Geochemists then use isotope signals preserved in biogenic materials to infer either what the biologic filter was (i.e., a species-specific biologic process), or the environment in which the animal lived (e.g., see review of Koch 1998). Although stable isotopes of hydrogen, carbon, nitrogen, oxygen, and strontium in modern-day animals have provided fruitful information on environment or biology, preservation of hydrogen...

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