Norman D. Smith, R. Stephen Saunders; Paleoenvironments and their control of acritarch distribution; Silurian of east-central Pennsylvania. Journal of Sedimentary Research ; 40 (1): 324–333. doi: https://doi.org/10.1306/74D71F37-2B21-11D7-8648000102C1865D
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Restricted to depositional areas open at least intermittently to marine waters, degree of preservation reflects depositional environment, distribution controlled by direction of prevailing currents at least in part
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