Two new species, Harpactocarcinus yozgatensis from the early Eocene (Ypresian) Yoncalı Formation of Turkey and H. dalmatius from the middle Eocene (Bartonian) Foraminiferal limestones of Hvar Island, Croatia, are described. Harpactocarcinus yozgatensis suggests an early Eocene origin of the genus. Their discovery posits a more easterly point of origin for the genus than was previously known, a position which is supported by the foraminiferal assemblages of both localities. Evaluation of species referred to Harpactocarcinus has resulted in one new combination, Neozanthopsis achalzicus (Bittner in Abich, 1882).

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