Scientific editing by Alistair Crame.

D. K. Loydell writes: In their recent paper Jeppsson & Aldridge (2000, p. 1137) expressed reservations about the method behind the generation of my (Loydell 1998) sea-level curve for the early Silurian (Llandovery to earliest Ludlow). As the method used was based upon decades of consistent observations of graptolite distribution, I am rather surprised that Jeppsson & Aldridge could not accept the scientific basis of the sea-level curve’s construction.

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Simply stated, graptolite diversity increases from low (usually 1–3 species) in nearshore environments to a maximum in outer shelf environments. Thus a significant rise in...

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