Upper Cenomanian peritidal limestones with dinosaur footprints are characterized by subtidal wackestone-packstone bearing rudist congregations as well as intertidal laminites. In the shallow subtidal environment radiolitids, as mud-supported elevator dwellers, thrived almost exclusively in dense thickets or floatstones with toppled individuals, that pass vertically and laterally into peloidal-foraminiferal wackestone-packstone. As is the case for the rudists, dinosaur tracks are also preserved within peritidal environments through the shallowing-upward cycles. However, unlike the rudists dinosaur tracks are present at the top of intertidal laminites. In cases when shallowing upward cycles ended with subtidal limestones, the possibility of dinosaur footprint preservation on the emerged surfaces was prevented by karstification of the upper bedding plane. Appearance of radiolitids through biostromal congregations, as well as preservation of dinosaurs footprints on their upper bedding plane, imply absence of possible barriers made by radiolitids on the vast shallow-water Adriatic carbonate platform during Upper Cenomanian.

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