Anatomy, Phylogeny and Palaeobiology of Early Archosaurs and their Kin

Archosaurs, an important reptile group that includes today’s crocodiles and birds, arose during the Triassic in the aftermath of the greatest mass extinction of all time. In the last 20 years, our understanding of the early evolution of the group has improved substantially with the discovery of new fossils and species of early archosaurs and their closest relatives, a better understanding of the relationships of these animals, and new insights into their palaeobiology. In order to synthesize these new data, researchers of early archosaurs from around the world met at the first symposium of early archosaur evolution at the IV Congreso Latinoamericano de Paleontología de Vertebrados (September 2011) in San Juan, Argentina. This symposium facilitated collaboration and strove to paint a better understanding of these extraordinary animals. The resultant body of work is a state-of-the-art examination of early archosaur groups and their close relatives including historical, anatomical, biogeographical, evolutionary and palaeobiological data. This contribution furthers our knowledge of the anatomy, relationships, and palaeobiology of species-level taxa as well as more global patterns of archosaur evolution during the Triassic.
Abstract
Pterosaurs are a clade of highly specialized, volant archosauromorphs recorded from the Upper Triassic to the uppermost Cretaceous. Problematic remains referred to the Pterosauria are reported from the Triassic of Europe and both North and South America, but unequivocal pterosaur specimens are only known from the Alps (Italy, Austria and Switzerland: Preondactylus buffarinii, Austriadactylus cristatus, Peteinosaurus zambellii, Eudimorphodon ranzii, Carniadactylus rosenfeldi, Caviramus schesaplanensis and Raeticodactylus filisurensis) and Greenland (‘Eudimorphodon’ cromptonellus). Pterosaurs are diagnosed mostly by features associated with the advent of powered flight. They are generally considered to be archosaurians more closely related to dinosaurs than to crocodilians, but non-archosaurian positions have also been proposed. There is a lack of general agreement about ingroup relationships, particularly among the basal pterosaurs. Triassic pterosaurs differ from other non-pterodactyloid pterosaurs in features of the dentition and caudal vertebral column. A ‘Big Bang’ model for their early history fits better with the fossil record: the earliest unequivocal pterosaurs show a sudden and geographically limited appearance in the fossil record, as well as a relatively high burst of diversity and considerable morphologic disparity. Absence of pterosaur remains from deposits where they are expected to be found suggests that they had not yet evolved in pre-Norian times.