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The description of a partial but well-preserved head of the sclerorhynchid batoid Sclerorhynchus atavus Woodward, 1889 gave the first clear indication of the presence of a puzzling group of extinct rostrum-bearing rays that resembled both the Pristidae (rays) and the Pristophoridae (sharks). Despite recognizing similarities to and differences from these extant groups, Smith Woodward suggested that Sclerorhynchus be assigned to the Pristidae, although acknowledging that the rostra are very different. Smith Woodward did note similarities of Sclerorhynchus rostrum saw-teeth to those of the Pristiophoridae, including the location of these along the margin of the rostrum, rather than in deep...

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