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An insect zonation with a time resolution of 1.5–2 Ma for Late Pennsylvanian to Early Permian (Kasimovian to Artinskian) non-marine deposits is presented. The zonation is based on the directed morphogenetic evolution of colour pattern in the forewings of the blattid (cockroach) family Spiloblattinidae. This evolution is observed in lineages of succeeding species of three genera. All three genera are widely distributed in the palaeo-equatorial zone from Europe to North America, that is, in the Euramerican biota province. Increasing reports of spiloblattinid zone species in condont-bearing, interfingered marine/continental strata of North American Appalachian, Mid-Continent and West Texas basins could...

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