The Johnnie Formation and associated Ediacaran strata in southwest Laurentia are ~3000 m thick, with a Marinoan cap carbonate sequence at the bottom, and a transition from Ediacaran to Cambrian fauna at the top. About halfway through the sequence, the Shuram negative carbon isotopic excursion occurs within the Rainstorm Member near the top of the Johnnie Formation, followed by a remarkable valley incision event. At its type locality in the northwest Spring Mountains, Nevada, the Johnnie lithostratigraphy consists of three distinctive sand-rich intervals alternating with four siltstone/carbonate-rich intervals, which appear correlative with other regional ­Johnnie Formation outcrops. Carbon isotope ratios in the sub–Rainstorm Member part of the Johnnie Formation are uniformly positive for at least 400 m below the Shuram excursion and compare well with sub–Shuram excursion profiles from the ­Khufai Formation in Oman. There is historical consensus that the Johnnie and overlying formations were deposited on a thermally subsiding passive margin. Following previous authors, we used Paleozoic horizons of known biostratigraphic age to define a time-dependent exponential sub­sidence model, and extrapolated the model back in time to estimate the ages of the Shuram excursion and other prominent Ediacaran horizons. The model suggests that the Shuram excursion occurred from 585 to 579 Ma, and that incision of the Rainstorm Member shelf occurred during the 579 Ma Gaskiers glaciation. It further suggests that the base of the Johnnie Formation is ca. 630 Ma, consistent with the underlying Noonday Formation representing a Marinoan cap carbonate sequence. Our results contrast with suggestions by previous workers that the Shuram excursion followed the Gaskiers event by some 20–30 m.y. We suggest instead that the Shuram and Gaskiers events were contemporaneous with the biostratigraphic transition from acantho­morphic to leiospherid acritarchs, and with the first appearance of widespread macroscopic animal life, 38 m.y. prior to the Ediacaran-Cambrian boundary.

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