Joseph J. Morley, Nicholas J. Shackleton; Extension of the radiolarian Stylatractus universus as a biostratigraphic datum to the Atlantic Ocean. Geology ; 6 (5): 309–311. doi: https://doi.org/10.1130/0091-7613(1978)6<309:EOTRSU>2.0.CO;2
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Comparison of the upper limit of Stylatractus universus with the δO18 record in two deep-sea cores from the mid- and low-latitude Atlantic reveals that this species becomes extinct within the transition zone between oxygen-isotope stages 12 and 11. This extends the global comparison of this biostratigraphic datum to the Atlantic, confirming previous studies that showed that the extinction of this species occurred on a world-wide basis approximately 425,000 yr ago.