Hayward (2001) drew attention to the little-known Pleistocene extinctions of deep-sea elongate benthic foraminifera and showed that the loss of diversity was far greater than previously reported. He concluded that these extinctions of elongate benthic taxa occurred during Pleistocene glacial periods, probably related to more vigorous circulation of cooler, more oxygenated bottom waters, and he suggested that the relationship between increased extinctions and global cooling was also apparent for two other major Cenozoic cooling periods, i.e., around the Eocene-Oligocene boundary and in the middle Miocene. Hayward's proposal is quite interesting, particularly in contrast to the recent studies of the mass...

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