Andreas Wetzel, Thomas Aigner; Stratigraphic completeness: Tiered trace fossils provide a measuring stick. Geology ; 14 (3): 234–237. doi: https://doi.org/10.1130/0091-7613(1986)14<234:SCTTFP>2.0.CO;2
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Trace-fossil associations commonly display a characteristic vertical zonation (tiering) within the substrate. When burrows of a particular tier become truncated by turbidites, tempestites, or other results of catastrophic events, they provide a simple but quantitative caliper for estimating stratigraphic incompleteness. The minimum amount of penecontemporaneous sea-floor erosion can be determined by types and sequences of trace fossils preserved below event deposits.