Sedimentation after wildfire is a profound disturbance to the biogeomorphic character of fluvial systems. Despite this significant alteration, field data focusing on the geomorphic processes and bed morphology that form the physical structure of habitat are limited, especially over a longer-term perspective. We report results of detailed field studies following two wildfires in the same steep fluvial system in the southern California Transverse Ranges three decades apart. Substantial channel sedimentation during storms following both fires altered physical elements of habitat, which included bed elevation, step height and spacing, pool depth and spacing, grain-size distribution (D₈₄), and transport capacity (τ_o/τ_c). The postfire sediment deposits buried bedforms, decreased D₈₄, and increased τ_o/τ_c by an order of magnitude. Incision during subsequent storms initiated recovery that depended on the attributes of post-wildfire climate variability that characterize the semiarid Mediterranean-type environment. Step-pool bedforms reappeared or reformed during the decades between wildfires as the relatively fine gravel-sized post-wildfire sediment was transported downstream, thus reestablishing the physical elements of habitat characterizing the dynamic system.