We have examined the deformation associated with a right-releasing stepover along the dextral Walker Lane belt where it traverses Wild Horse Mesa in eastern California. We use a micropolar inversion of both seismic focal mechanism and fault-slickenline data and compare the results to the micropolar deformation parameters inferred from paleomagnetically determined block rotations and GPS velocities. The focal mechanisms, fault-slickenlines, and GPS velocities all show horizontal shear with a consistent ENE–WSW to E–W maximum extension-rate axis (d ₁). A subset of data shows crustal thinning with a similarly oriented d ₁. We interpret these results as a reflection of divergent strike-slip (i.e., transtensional) boundary conditions in a negative flower structure developed in the right-releasing stepover. The fault-slickenline data also show a crustal thickening solution that we attribute to the local accommodation of block rotations. Paleomagnetic data demonstrate clockwise-looking-down rotations of 12.0° ± 2.6° (68% confidence limits) in ca. 3 Ma volcanic rocks, relative to the same rocks outside the stepover. Assuming rotations took 2–3 m.y. gives average microspins (block rotation rates) of 4.0° ± 0.9°/m.y. to 6.0° ± 1.3°/m.y. GPS velocities define a current macrospin (half the continuum rotation rate) of 3.9° ± 0.6°/m.y. to 6.1° ± 1.5°/m.y. These spin components are consistent with expectations for transtension. Our calculations of relative vorticity W from the GPS and paleomagnetic data are generally consistent with values obtained from the inversion of the fault-slickenline data, but the uncertainties in the data do not permit a definitive test of these results.