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The Late Devonian Red Hill locality in north-central Pennsylvania contains an Archaeopteris-dominated plant fossil assemblage, a diverse fossil fauna, and an extensive sedimentary sequence ideal for investigating the landscapes and biotic associations of the earliest forest ecosystems. Sedimentological analysis of the main plant-fossil bearing layer at Red Hill indicates that it was a flood-plain pond. A seasonal wet-and-dry climate is indicated by well-developed paleovertisols. The presence of charcoal interspersed with plant fossils indicates that fires occurred in this landscape. Fires appear to have primarily affected the fern Rhacophyton. The specificity of the fires, the distribution profile of the...

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