Recognition of differences in the habitats, apparatuses, and ranges of Late Devonian Icriodus and Pelekysgnathus permits refinement of their biofacies interpretations and construction of an alternate icriodontid zonation. Icriodus is a euphotic genus that predominated in most environments during the early Late Devonian (Frasnian) but died out during the early Famennian. Its apparatus consists of platform (I) elements; four larger, acodiniform cones; and two smaller, oneotodiform, scolopodiform, or drepanodiform cones. Pelekysgnathus is a shallow-water genus, which shortly after Icriodus died out, produced somewhat deeper water taxa with triple-rowed I elements that are homeomorphs of Icriodus I elements. Apparatuses for both...