Abstract Effective radium-226 concentration, EC Ra , is the product of radium activity concentration, C Ra , multiplied by the emanation coefficient, E , which is probability of producing a radon-222 atom in the pore spaces. It is measured by accumulation experiments in the laboratory, achieved routinely for a sample mass >50 g using scintillation flasks to measure the radon concentration. We report on 3370 EC Ra values obtained from more than 11 800 such experiments. Rocks ( n =1351) have a mean EC Ra value of 1.9±0.1 Bq kg −1 (90% of data in the range 0.11–35 Bq kg −1 ), while soils ( n =1524) have a mean EC Ra value of 7.5±0.2 Bq kg −1 (90% of data between 1.4 and 28 Bq kg −1 ). Using this large dataset, we establish that the spatial structure of EC Ra is meaningful in geology or sedimentology. For plants ( n =85), EC Ra is generally <1 Bq kg −1 , but values of larger than 10 Bq kg −1 are also observed. Dedicated experiments were performed to measure emanation, E , in plants, and we obtained values of 0.86±0.04 compared with 0.24±0.04 for sands, which leads to estimates of the radium-226 soil-to-plant transfer ratio. For most measured animal bones ( n =26), EC Ra is >1 Bq kg −1 . Therefore, EC Ra appears essential for radon modelling, health hazard assessment and also in evaluating the transfer of radium-226 to the biosphere.

Abstract: After being inaccessible for a number of years, the holotype and other specimens of the dsungaripterid pterodactyloid pterosaur Noripterus complicidens are again available for study. Numerous taxa assigned to the Dsungaripteridae have been described since the erection of Noripterus , but with limited comparisons to this genus. Based on the information from Young’s original material here we revise the taxonomic identity of N. complicidens and that of other Asian dsungaripterids. We conclude that N. complicidens is likely to be distinct from the material recovered from Mongolia and this latter material should be placed in a separate genus.

Abstract: A specimen of a pterodactyloid pterosaur from the Upper Tunbridge Wells Sand Formation (Early Cretaceous, Valanginian) of Bexhill, East Sussex, southern England is described. It comprises a small fragment of jaw with teeth, a partial vertebral column and associated incomplete wing bones. The juxtaposition of the bones suggests that the specimen was originally more complete and articulated. Its precise phylogenetic relationships are uncertain but it represents an indeterminate lonchodectid with affinities to Lonchodectes sagittirostris ( Owen 1874 ) which is reviewed here, and may belong in Lonchodraco Rodrigues & Kellner 2013 . This specimen is only the third record of pterosaurs from this formation.

Abstract: The Middle–Upper Jurassic Oxford Clay Formation of the UK is a highly productive fossiliferous unit, containing numerous fossils of marine and terrestrial vertebrates. Despite this abundance, pterosaurs make up only a tiny percentage of the assemblage. There are currently only seven specimens accessioned in the Oxford University Museum of Natural History and Natural History Museum, London. Of these, only one specimen (an association of material from a juvenile animal) has been named, identified as Rhamphorhynchus jessoni . Following a detailed examination, it is identified here as lacking any autapomorphies and the species R . jessoni is considered a nomen dubium . The other Oxford Clay Formation pterosaur fossils are similarly identified as indeterminate, except for an isolated scapulocoracoid which is positively identified as Rhamphorhynchus sp. The rarity of pterosaurs is attributed to a combination of the fragility of their bones and the tendency of volant animals to have low population densities at sea.

Abstract: An incomplete bone from the latest Cretaceous dinosaur site of Villaggio del Pescatore (Trieste Province, Italy) is definitely a wing metacarpal of a pterodactyloid pterosaur. It represents the only Italian Cretaceous pterosaur remains known, as well as the only pterosaur from the Adriatic Carbonate Platform. With an estimated minimum length of 136 mm, it belongs to a relatively small individual relative to the standard of latest Cretaceous pterodactyloids. It is not as elongated and gracile as azhdarchid wing metacarpals and shows a mix of features found in Pteranodon and some more basal pterodactyloids. It is one of the very few remains of putative non-azhdarchid pterosaurs from the upper Campanian–Maastrichtian worldwide and supports the view that the Azhdarchidae were not the only pterosaur clade existing during latest Cretaceous times.

Abstract: The Solnhofen pterosaurs Pterodactylus antiquus , Aerodactylus scolopaciceps , Diopecephalus kochi , Germanodactylus cristatus and Germanodactylus rhamphastinus all have complicated taxonomic histories. Species originally placed in the genus Pterodactylus , such as Aerodactylus scolopaciceps , Ardeadactylus longicollum , Cycnorhamphus suevicus and Germanodactylus cristatus possess apomorphies not observed in the type species of Pterodactylus , and consequently have been placed in new genera. The affinities of another Solnhofen pterosaur previously placed in Pterodactylus , Diopecephalus kochi , are less clear. It has been proposed that D. kochi is a juvenile specimen of Pterodactylus antiquus , or perhaps ‘ Germanodactylus rhamphastinus ’ specimens are mature examples of D. kochi . Furthermore, studies have suggested that ‘ Germanodactylus rhamphastinus ’ is not congeneric with the type species of Germanodactylus. Geometric morphometric analysis of prepubes and a cladistic analysis of the Pterosauria elucidate plesiomorphic and apomorphic conditions for basal Jurassic pterodactyloids. Germanodactylus is found to be a monotypic genus and Pterodactylus , Diopecephalus , and ‘ G. rhamphastinus ’ are found as distinct taxa belonging in individual genera, diagnosable using a combination of characters. Thus, Diopecephalus kochi is not demonstrated to be congeneric with Germanodactylus or Pterodactylus and is maintained as a valid taxon. ‘ G. rhamphastinus ’ is readily distinguishable from other Solnhofen pterosaur taxa, and a new genus is erected for its reception. Supplementary material: Technical details of the mechanical theory behind this article are available at https://doi.org/10.6084/m9.figshare.c.3809401.v1

Abstract: Pterosaur fossils are rare in the Morrison Formation, and most are poorly preserved. The Breakfast Bench Facies (BBF) at Como Bluff produces incomplete but uncrushed limbs. One proximal and two distal femora match a complete femur (BYU 17214) referred to Mesadactylus . Unexpectedly, both of the BBF distal femora possess a large intercondylar pneumatopore. BYU 17214 also possesses an intercondylar pneumatopore, but it is smaller than in the BBF femora. Distal femoral pnuematicity is previously recognized only in Cretaceous azhdarchoids and pteranodontids. A peculiar BBF jaw fragment shows strongly labiolingually compressed, incurved crowns with their upper half bent backwards; associated are anterior fangs. We suspect this specimen is a previously undiagnosed pterosaur. Additional BBF material documents a diverse pterosaur fauna including a humerus with a greatly expanded ectepicondyle possibly from a non-pterodactyloid monofenestratan.

Abstract: Six specimens accessioned to the Bavarian State Collection for Palaeontology and Geology in Munich, Germany, in 1966 are identified as coming from a gigantic pterodactyloid pterosaur. The previously undescribed material was obtained in 1955 by Jean Otto Haas and compares favourably in size with the type specimen of the Late Cretaceous (Maastrichtian) azhdarchid pterosaur Arambourgiania philadelphiae ( Arambourg 1959 ) from the same locality/region. The material represents fragments of two cervical vertebrae, a neural arch, a left femur, a ?radius, and a metacarpal IV and bones of problematic identity, and does not duplicate the type material of Arambourgiania . The timing of its collection and its locality of Ruseifa, Jordan suggest it might pertain to the same individual as the holotype.

Abstract: Three associated incomplete wing bones of a large pterodactyloid pterosaur from the Ferron Sandstone Member of the Mancos Shale Formation of Utah are described. Based on the morphology of the bones’ articular ends, the specimen is referred to the Pteranodontoidea. Associated ammonites indicate that the specimen’s age is Middle Turonian, so it adds to the sparse worldwide record of Turonian pterosaurs. The record of pteranodontoid pterosaurs in the Western Interior Seaway of North America is reviewed, and it is suggested that pteranodontoids were not diverse.

Abstract: A pterosaur humerus (PRC 64) from the Upper Jurassic of Thailand was initially assigned to the Azhdarchoidea, an important clade of edentulous Cretaceous pterodactyloid pterosaurs. Here it is reassigned, on the basis of morphological comparisons and a phylogenetic analysis, to the Rhamphorhynchidae, a widely distributed clade of small to medium-sized seemingly piscivorous basal pterosaurs. This record extends the geographical range of rhamphorhynchids and confirms recent findings that these pterosaurs inhabited continental as well as marginal marine ecosystems.