Abstract The Paleozoic Variscan orogen in Europe has a markedly circuitous trace for which several different origins have been postulated, including deformation around promontories on the colliding continental margins, extrusion within the collision zone, folding of a ribbon continent and collision with a substantial dextral component. Adopting the latter assumption, unfolding of the large, steeply plunging folds (‘oroclines’) of Iberia and the Moroccan Meseta requires more than 4000 km of dextral lateral translation of Laurussia with respect to Gondwana. Constraints on the age of the folding require that this lateral translation occurred in mid-late Carboniferous time. Significant dextral translation of Laurussia with respect to Gondwana late in the Variscan collision is supported by palaeomagnetic data for the two supercontinents, although the exact timing of this relative motion is not well constrained due to large uncertainties in the palaeomagnetic pole positions. Our palinspastic reconstruction of the major Variscan folds of Iberia places the rocks of the South Portuguese Zone of western Iberia adjacent to the Rhenish Massif; for instance, as part of Avalonia. By the same token, the Sehoul Block in the Moroccan Meseta probably originated at the western end of Cadomia, although nothing in our analysis precludes it also being derived from Avalonia.

Abstract The extensive, predominantly siliciclastic deposits of the Upper Ordovician of the Tafilalt have long been the subject of scientific investigation. In the past 25 years, intensified collecting for commercial purposes has resulted in the discovery of several exceptionally-preserved faunas (Konservat-Lagerstätten) in the Tafilalt region, preserving a range of non-biomineralized and soft-bodied organisms. The preservation of these fossils in the coarse clastic sediments of the Tafilalt is surprising, and in the case of soft-bodied organisms, remarkably similar to the preservational mode of typical Ediacaran biotas. These relatively recent discoveries have increased the scientific significance of the Tafilalt Biota, providing an unparalleled insight into the composition and temporal evolution of the shallow, open-marine ecosystems and their denizens during the later stages of the Great Ordovician Biodiversification Event. At least nine different phyla, in addition to several soft-bodied problematica are represented in the Tafilalt. While the highly diverse and remarkably well-preserved echinoderm and euarthropod faunas are most emblematic for the Tafilalt Biota, further studies have revealed a relatively high diversity of molluscs and brachiopods. Among soft-bodied fossils, the problematic paropsonemid eldonids are iconic for the Tafilalt and stand out both through their abundance, and their wide temporal and geographical range throughout the area.