ABSTRACT During the Pleistocene, the Laurentian Ice Sheet extended southward into northwestern Pennsylvania. This field trip identifies a number of periglacial features from the Appalachian Plateaus and Ridge and Valley provinces that formed near the Pleistocene ice sheet front. Evidence of Pleistocene periglacial climate in this area includes glacial lake deposits in the Monongahela River valley near Morgantown, West Virginia, and Sphagnum peatlands, rock cities, and patterned ground in plateau areas surrounding the Upper Youghiogheny River basin in Garrett County, Maryland, and the Laurel Highlands of Somerset County, Pennsylvania, USA. In the high-lying basins of the Allegheny Mountains, Pleistocene peatlands still harbor species characteristic of more northerly latitudes due to local frost pocket conditions. Pleistocene fauna preserved in a cave deposit in Allegany County, Maryland, record a diverse mammalian assemblage indicative of taiga forest habitat in the Ridge and Valley province.

ABSTRACT With waterfalls and the deepest gorge in Pennsylvania, Ohiopyle State Park provides opportunities to observe a variety of habitats and three-dimensional (3-D) exposures of the Pennsylvanian sandstone most responsible for shaping Laurel Highlands landscapes. Evidence for the relationship between bedrock, ancient climates, and the landscape can be observed at some of the most scenic natural features of the park: Baughman Rock Overlook, Cucumber Falls, Ohiopyle Falls, Meadow Run Waterslide and Cascades, and Youghiogheny River Entrance Rapid. Channel azimuths and lateral variations in thickness of upper Pottsville fluvial/deltaic sandstone suggest that deposition was influenced by deformation of this part of the Allegheny Plateau during the Alleghanian orogeny. Geologic features of Pottsville sandstone outcrops include a 10-m- (~33-ft-) long Lepidodendron fossil and a 3-D exposure of a meter-high Pennsylvanian subaqueous sand dune and scour pit. Cosmogenic age dating has indicated very slow erosion of hard sandstone in an upland location at Turtlehead Rock and informed estimation of Pleistocene/Holocene waterfall retreat rates of Ohiopyle and Cucumber Falls. Bedrock exposures supporting scour habitats along the Youghiogheny River occur only in a limited area of Youghiogheny Gorge where knickpoint migration and bedrock erosion were relatively recent. Geologic factors, including locations of major tributaries, development of bars that constrict river flow, and proximity of Homewood sandstone outcrops as sources of boulder obstacles in the river, contributed to the class, location, and nature of whitewater rapids in the lower Youghiogheny River.

Abstract The Late Pennsylvanian was a time of ice ages and climate dynamics that drove biotic changes in the marine and non-marine realms. The apex of late Paleozoic glaciation in southern Gondwana was during the Late Pennsylvanian, rather than the early Permian as inferred from more equatorial Pangaea. Waxing and waning of ice sheets drove cyclothemic sedimentation in the Pangaean tropics, providing an astrochronology tuned to Earth-orbital cycles, tied to climatic changes, reflected in aeolian loess and palaeosol archives. Vegetation change across the Middle–Late Pennsylvanian boundary was not a ‘Carboniferous rainforest collapse’, but instead a complex and drawn out step-wise change from one kind of rainforest to another. Changes in marine invertebrate and terrestrial vertebrate animals occurred across the Middle–Late Pennsylvanian boundary, but these did not lead to substantive changes in the organization of those communities. The base of the Upper Pennsylvanian is the base of the Kasimovian Stage, and this boundary needs a GSSP to standardize and stabilize chronostratigraphic usage. To avoid further chronostratigraphic confusion, the Cantabrian Substage should be abandoned, and the traditional Westphalian–Stephanian boundary should be returned to and recognized as the time of major floristic change, the lycospore extinction event.

Abstract The Kasimovian Stage is the lower stage of the Upper Pennsylvanian Subsystem, in which a series of considerable biotic and abiotic events happened and changed the Earth. The Variscan orogeny and the Late Paleozoic Glaciation are two major events that caused geographical isolation of marine faunas and difficulties for a global correlation of biostratigraphy. Regional timescales of the Kasimovian across major continents are reviewed here. A global correlation of the Kasimovian is tentatively established based on a detailed review of major fossil groups such as conodonts, fusulines and some macrofossils. The index taxon for the base of the Kasimovian Stage has not been selected. The conodont species Swadelina subexcelsa , Idiognathodus heckeli , I. turbatus and I. sagittalis have good potential. Among them, I . heckeli is considered the best marker for the base of the Kasimovian Stage because it can mark a bioevent in a wide geographical range, and more importantly, it has a clear taxonomic definition within a phylogenetic lineage. The fusuline Montiparus might be regarded as an auxiliary marker to define the base of the Kasimovian based on its wider distribution. Other proxies, i.e. isotopic dating and strontium, carbon and oxygen isotopic stratigraphy throughout the Kasimovian, are also reviewed. The Global Boundary Stratotype Section and Point (GSSP) candidates for the Kasimovian Stage include the Naqing section, South China, the Usolka section, South Urals and the Afanasievo section, Moscow Basin. The Naqing section is regarded as the most appropriate GSSP candidate in terms of its complete sedimentary succession, well-recorded conodont lineages and well-studied bio-, chemo- and cyclo-stratigraphy.

Abstract For the west European regional chronostratigraphic framework, the Cantabrian substage was conceived as covering a widely apparent stratigraphic gap between the top of the Westphalian and the base of Stephanian A, the lowest unit of the Stephanian. A continuous depositional history covers this time gap in the Cantabrian region of Spain; the upper limit of this interval was defined by the succeeding Barruelian substage, equivalent to Stephanian A. Intense tectonic and magmatic activity characterizes this period; the Iberian orogenic belt was an essentially linear feature buckled through the Late Pennsylvanian into the tightly folded Cantabrian Orocline. This evidences an extensive southern foreland to the Variscides, in which the coal-swamp biome persisted through the Late Pennsylvanian, supporting biostratigraphical correlation with the Donbass. New high precision U–Pb CA-ID-TIMS radiometric dating of tonstein horizons supports a preliminary time-framework of regional substages: base of the Asturian (proposed, ex-Westphalian D) c. 310.7 Ma; base of the Cantabrian c. 307.5 Ma; base of the Barruelian (ex-Stephanian A) c. 304.9 Ma; base of the Saberian (proposed) c. 303.5 Ma. The Cantabrian and Barruelian embrace the entire Kasimovian of the global time-scale, and the top of the Barruelian is essentially coincident with the base of the Gzhelian.

Abstract In spite of numerous revisions from 1966 to present, the Cantabrian Substage of the Stephanian Stage (Pennsylvanian) was never properly defined as a chronostratigraphic unit. Defined and redefined at least three times, the Cantabrian lacks boundary stratotypes that correspond to clear and correlateable biochronological signals. Thus, instead of using a biochronological datum of well-established validity and utility, Cantabrian advocates have relied on ill-defined macrofloral assemblage zones and on lithostratigraphic boundaries to define the substage. As a result, the Cantabrian is demonstrably diachronous, even within Europe; indeed, the Cantabrian has proven to be unusable for correlations outside its type area in northern Spain. To resolve these problems, we recommend that the Cantabrian Substage be abandoned, and the Westphalian–Stephanian boundary be redefined at the major floral turnover that has been documented in the USA, western and central Europe, and in the Donets Basin. We further recommend that the bases of the Kasimovian Series, Stephanian Series, Missourian Series, and Upper Pennsylvanian Series all be aligned with this same floral turnover.

Abstract The Kasimovian was a time of ecological upheaval and large-magnitude changes in palaeoclimate. Referred to as the ‘collapse’ of the palaeotropical rainforests, the Kasimovian is marked by rapid changes in megafloral communities and associated ecosystem effects on vertebrates and invertebrates. p CO 2 variation coincided with these ecological catastrophes, varying between pre-industrial levels (PAL) to 2×PAL on 10 5 year timescales. Our understanding of the carbon cycle perturbations that affected p CO 2 and the connection of these climate-forcings to the terrestrial upheaval of palaeotropical rainforests remains a grand challenge. Here, the effects of palaeosol accumulation and/or degradation on the terrestrial carbon cycle during the Kasimovian is assessed. Palaeosols are surveyed from ice-free depositional basins on Pangaea and assessed for palaeolandscape equilibrium. An orbital framework is developed in order to understand the relationships of palaeosols, the carbon cycle, and insolation. Based on these analyses a key time interval emerges in the early Kasimovian. This time interval records a shift in palaeolandscape equilibria, terrestrial carbon cycling, and orbital forcing. The carbon cycling and landscape equilibria are eccentricity-paced; however, predominance of short eccentricity and obliquity throughout this interval indicates that the changes to palaeosols and the locus of carbon burial may have acted as a stochastic process.

Abstract In Europe, North Africa and Asia Minor, the remains of Pennsylvanian sedimentary basins bearing continental deposits either intimately mixed with shallow-marine strata or deposited in exclusively continental settings are preserved. Long-lasting research on these basins allowed the definition of regional stages and substages based on marine fauna and terrestrial flora, later extended by terrestrial and freshwater faunal biostratigraphies. Glacioeustatically driven marine bands provide laterally widespread correlation markers; however, where such bands are missing only biostratigraphic control exists. Resolution of biostratigraphic zonations combined with gaps in sedimentary successions and variable quality of the fossil record throughout the basin fills do not allow in all cases a precise correlation between the Pennsylvanian basins in Europe and, in turn, the timing of tectonic, climatic and biotic events, and thus an absolute complete understanding of the response of terrestrial and freshwater biota to climate changes across eastern tropical Pangaea. A helpful tool is new radioisotopic ages of intercalated volcaniclastics that reveal the partial diachroneity of some widely used biostratigraphies. We attempt to present the current state of the art to stimulate further research to mitigate gaps in our knowledge.

Abstract A threshold-like vegetational change in tropical wetlands occurred in the early Kasimovian (the US Desmoinesian–Missourian boundary) – Event 3. Two earlier significant changes occurred, first in the mid-Moscovian (Atokan–Desmoinesian; ∼Bolsovian–Asturian) – Event 1, and the second in the late Moscovian (mid-Desmoinesian; mid-Asturian) – Event 2. These changes occurred during a time period of dynamic and complex physical change in Euramerican Pangaea driven by changes in polar ice volume and accompanying changes in sea level, atmospheric circulation, rainfall, and temperature. During the Event 3 change, hyperbolized as ‘the Carboniferous rainforest collapse’, lycopsid dominance of (mostly peat) swamps changed to marattialean tree-fern and medullosan pteridosperm dominance, and biodiversity decreased. Event 3 encompassed one glacial–interglacial cycle and included vegetational turnover in other wetland habitats. For several subsequent glacial–interglacial cycles peatland dominance varied, known from palynology, before stabilizing. These vegetational changes likely reflect climatic events driving unidirectional, non-reversible wetland vegetational changes, during cooler, wetter parts of glacial–interglacial cycles. Discussion is complicated by different placements of crucial stratigraphic boundaries, but under the same names, compromising both clear communication and understanding of the literature. Not the least is the floating base of the Cantabrian Substage, together with the position of the Westphalian–Stephanian Stage boundary.

Abstract We present the first analysis of vegetational change in far western equatorial Pangaea (New Mexico, USA) during the Middle–Late Pennsylvanian transition (determined by conodonts and fusulinids) of the Late Paleozoic Ice Age. The study is based on the largest database assembled from this region: 28 of 44 quantitatively analysed floras from 14 of 26 stratigraphic levels. Most sampled floras are ‘mixed’, both below and above the boundary, including both hygromorphic and mesomorphic/xeromorphic taxa. The taxonomic data were recalibrated morphometrically focusing on foliar traits of lamina width and venation. All data were examined using stratigraphic credible intervals, capture–mark–recapture analyses, and resampling analyses. Results indicate no substantive taxonomic turnover across the boundary. This stands in marked contrast to patterns in mid-Pangaean coal basins where there is a large wetland vegetational turnover. However, plant and physical geological data indicate that immediately following the boundary in New Mexico, and for approximately half of the Missourian Stage, floras previously dominated by hygromorphs become overwhelmingly dominated by mesomorphic/xeromorphic taxa. Although expressed differently, the western Pangaean physical and palaeobotanical patterns parallel those from mid-Pangaean coal basins and suggest a widespread environmental change.

Abstract A series of vegetation changes take place in tropical ecosystems during the Pennsylvanian Subperiod. The most notable change, recognizable from palynology and plant macrofossils at the Middle and Late Pennsylvanian boundary in the Illinois Basin, is the extirpation, or local extinction, of certain lineages of arborescent lycopsids, followed by their replacement by stem group marattialean tree ferns. The leading hypothesis suggests a significant change in precipitation regime as the cause. To test this hypothesis, we examine the vascular anatomy and physiology of key lineages of Pennsylvanian plants: the sphenopsids, tree ferns, cordaitaleans, medullosans, lycophytes and extrabasinal stem group coniferophytes. Using scanning electron and light microscopy of fossilized anatomy, we provide new data on these plants’ vascular systems, quantifying their physiological capacity and drought resistance. We find that three Pennsylvanian plant lineages – the medullosans, arborescent lycopsids and Sphenophyllum – contain high hydraulic conductivity but are vulnerable to drought-induced damage, whereas others are resistant, including stem group tree ferns and coniferophytes. Relative abundance changes among these plants were likely driven by drought, and differences in water use efficiency would have amplified drought events as plant communities changed. The interaction of physiological selectivity and positive feedback between aridity and drought tolerance likely played a significant role in Late Paleozoic floral changes.

Abstract Late Pennsylvanian conodont faunas were dominated by idiognathodids historically assigned to Idiognathodus (flat P 1 ) or Streptognathodus (troughed P 1 ). Recent work suggests clades arose iteratively, through time, from unrelated ancestors in different geographical regions. The end-Desmoinesian extinction event terminated two major genera, Swadelina (troughed) and Neognathodus (long carina), and comparable new morphotypes developed from surviving Idiognathodus species in the early Kasimovian, especially in North America. True Streptognathodus (troughed) and Heckelina n. gen (asymmetric, eccentric groove) appeared in North America in the mid-Kasimovian. Another troughed clade arose in Eurasia (‘ S. ’ 2) and attained a global distribution by the late Kasimovian. A second, early Gzhelian, Eurasian radiation produced new troughed forms (‘ S. ’ 4) that dominated Gzhelian faunas globally. In South China, endemic clades of eccentrically grooved Idiognathodus ? and troughed forms (‘ S .’ 3) appeared in the late Kasimovian and persisted into the Gzhelian. Typical Idiognathodus species were uncommon by the late Kasimovian and disappeared in the mid-Gzhelian. After a low diversity interval in the mid-Gzhelian, a new major radiation of weakly troughed forms occurred (‘ S. ’ 5), which led to redevelopment of Idiognathodus -like elements in the Cisuralian. Other conodont genera from offshore ( Gondolella, Idioprioniodus ) and nearshore settings ( Hindeodus, Diplognathodus, Adetognathus, Ellisonia ) are poorly studied and show low diversity and little morphological change.

Abstract Late Moscovian–early Gzhelian conodonts occur abundantly in a newly discovered slope section, the Shanglong section, southern Guizhou, South China. The conodont fauna is dominated by P1 elements of Idiognathodus and associated with elements of Swadelina , Streptognathodus and Heckelina . A total of 62 species, including species in open nomenclature, were identified, which are assigned to eight genera. Index conodont species of Middle and Late Pennsylvanian, e.g. I. podolskensis Group, Sw. sp. A, Sw . subexcelsa , Sw . makhlinae , I . heckeli , I . magnificus , I . guizhouensis , H. eudoraensis , I . naraoensis , and H . simulator are all recovered, and their 10 conodont zones are recognized. The richness and abundance of the conodonts throughout the section are analysed. Conodont richness ranges from 1 to 14 and is positively related to conodont abundance (1–379). The composition of conodont elements, i.e. sinistral v. dextral, P1 v. non-P1 and adult v. subadult and juvenile, is presented. The numerical cluster technique is employed to identify four subbiofacies of the slope setting, namely the I . podolskensis , Swadelina , I. swadei–magnificus and Streptognathodus – Heckelina–Idiognathodus subbiofacies.