Petrologic, geochemical, and metamorphic data on gneissic xenoliths derived from the middle and lower crust in the Neogene Bering Sea basalt province, coupled with U-Pb geochronology of their zircons using sensitive high-resolution ion microprobe–reverse geometry (SHRIMP-RG), yield a detailed comparison between the P-T-t and magmatic history of the lower crust and magmatic, metamorphic, and deformational history of the upper crust. Our results provide unique insights into the nature of lithospheric processes that accompany the extension of continental crust. The gneissic, mostly mafic xenoliths (constituting less than two percent of the total xenolith population) from lavas in the Enmelen, RU, St. Lawrence, Nunivak, and Seward Peninsula fields most likely originated through magmatic fractionation processes with continued residence at granulite-facies conditions. Zircon single-grain ages (n = 125) are interpreted as both magmatic and metamorphic and are entirely Cretaceous to Paleocene in age (ca. 138–60 Ma). Their age distributions correspond to the main ages of magmatism in two belts of supracrustal volcanic and plutonic rocks in the Bering Sea region. Oscillatory-zoned igneous zircons, Late Cretaceous to Paleocene metamorphic zircons and overgrowths, and lack of any older inheritance in zircons from the xenoliths provide strong evidence for juvenile addition of material to the crust at this time. Surface exposures of Precambrian and Paleozoic rocks locally reached upper amphibolite-facies (sillimanite grade) to granulite-facies conditions within a series of extension-related metamorphic culminations or gneiss domes, which developed within the Cretaceous magmatic belt. Metamorphic gradients and inferred geotherms (~30–50 °C/km) from both the gneiss domes and xenoliths are too high to be explained by crustal thickening alone. Magmatic heat input from the mantle is necessary to explain both the petrology of the magmas and elevated metamorphic temperatures. Deep-crustal seismic-reflection and refraction data reveal a 30–35-km-thick crust, a sharp Moho and reflective lower and middle crust. Velocities do not support a largely mafic (underplated) lower crust, but together with xenolith data suggest that Late Cretaceous to early Paleocene mafic intrusions are likely increasingly important with depth in the crust and that the elevated temperatures during granulite-facies metamorphism led to large-scale flow of crustal rocks to produce gneiss domes and the observed subhorizontal reflectivity of the crust. This unique combined data set for the Bering Shelf region provides compelling evidence for the complete reconstitution/re-equilibration of continental crust from the bottom up during mantle-driven magmatic events associated with crustal extension. Thus, despite Precambrian and Paleozoic rocks at the surface and Alaska’s accretionary tectonic history, it is likely that a significant portion of the Bering Sea region lower crust is much younger and related to post-accretionary tectonic and magmatic events.

The Kodiak Border Ranges ultramafic complex, Afognak batholith, and Shuyak Formation on Kodiak and Afognak Islands together form the lower, middle, and upper portions, respectively, of a Jurassic–Triassic island-arc crustal section. The Kodiak section exhibits structural and geochemical trends similar, but not identical to, the Tonsina-Nelchina segment of the Talkeetna arc, located >500 km to the northeast. Exposed at the base of the Kodiak section is cumulate clinopyroxenite with associated dunite, wehrlite, and layered gabbro. In the inferred middle to upper crust, tonalite and quartz diorite of the Afognak batholith intrude Shuyak Formation basaltic flows, basaltic pillow lavas, and volcaniclastic sedimentary rocks. Despite the fault-bounded nature of the lower crustal and mantle rocks, continuous chemical trends in elements such as MgO, Ni, Cr, Nb, Sr, Y, and rare-earth elements exist across all three units. Modeling of these data suggest that Kodiak arc evolution occurred in two main stages: (1) a gabbroic initial melt underwent fractional crystallization that produced a pyroxenitic root and a gabbroic lower crust, and (2) melt in equilibrium with the gabbroic lower crust underwent assimilation-fractional crystallization to produce mid-crustal plutonic and upper-crustal volcanic rocks. Kodiak Island exposes the oldest and thinnest portion of the Talkeetna arc, with ages from the Afognak batholith ranging from ca. 215–185 Ma. In the eastern and western Talkeetna arc, magmatism migrated northward after ca. 180 Ma in response to inferred forearc erosion. Forearc erosion coupled with differential subduction-channel movement juxtaposed blueschist-facies rocks with middle and lower crustal arc rocks. These processes occurred earlier and to a greater degree in the western Talkeetna arc, causing the arc to split in half, separating the Kodiak and Alaskan Peninsula parts of the Talkeetna arc.

Fossils within accreted terranes are typically used to describe the age or origin of the exotic geologic blocks. However, accretion may also provide new pathways for faunal exchange between previously disconnected landmasses. One such landmass, the result of accretion, is Beringia, that entity encompassing northeastern Asia and northwestern North America and the surmised land connection between the two regions. The present concept of Beringia as a Quaternary subcontinent includes a climatic component in the form of glacial advances and retreats driving changes in sea level. These changes may have facilitated exchanges of marine biota between the Pacific Ocean and Arctic Basin, or exchanges of terrestrial faunas and floras between Asia and North America. The Beringian ecosystem includes specializations of the flora and fauna, especially in the vertebrate fauna. A review of tectonic reconstructions and the striking taxon-free parallel patterns in data on the Cretaceous and Quaternary fauna and flora suggest that a generalized concept of Beringia should be formally extended back in time to the Cretaceous. A significant shift in emphasis of defining variables occurs with this extension. Climate, in the form of meteorological phenomena, and geologic history are important variables in the previously recognized definition of Beringia. The extension of Beringia into the Cretaceous implies that Beringia is rooted in its accretionary rather than its climatic history; in other words, the geographic pattern as the result of tectonics is the defining parameter for Beringia.