Abstract Upper Carboniferous stages in terrestrial strata are consistently recognizable throughout the Euramerican Realm based on the ranges of plant macrofossils rather than named biozones. Uniformitarianism is an invaluable principle used to understand much of Earth's history. However, it has been recognized that relatively short time intervals exist when major changes occurred in the biota and sedimentation style at a non-uniformitarian time scale. Many well-defined Upper Carboniferous stage boundaries are located at such events. The Cantabrian Stage was proposed in 1969 based on the assumption that the ‘Florensprung’ (‘floral jump’) of Gothan, a dramatic change in terrestrial floras at the traditional Westphalian–Stephanian boundary, indicated that strata were missing, and the postulated ‘gap’ had to be closed by finding strata representing this ‘missing’ time elsewhere. The suggestion of a ‘gap’ or hiatus at this level reflects uniformitarian thinking, but is incorrect. As recent work has shown that a drastic climatic change reached a threshold in the tropical palaeoequatorial parts of Pangaea at the Westphalian–Stephanian (approximately the Moscovian–Kasimovian) boundary as a consequence of a drying trend, floras changed rapidly, and the traditional Westphalian–Stephanian boundary is thus one of the most easily recognizable biostratigraphic boundaries in Pennsylvanian terrestrial beds. The Cantabrian Stage, proposed to fill the non-existent gap, does not exist and cannot be recognized, either in its type area or elsewhere.

Abstract A threshold-like vegetational change in tropical wetlands occurred in the early Kasimovian (the US Desmoinesian–Missourian boundary) – Event 3. Two earlier significant changes occurred, first in the mid-Moscovian (Atokan–Desmoinesian; ∼Bolsovian–Asturian) – Event 1, and the second in the late Moscovian (mid-Desmoinesian; mid-Asturian) – Event 2. These changes occurred during a time period of dynamic and complex physical change in Euramerican Pangaea driven by changes in polar ice volume and accompanying changes in sea level, atmospheric circulation, rainfall, and temperature. During the Event 3 change, hyperbolized as ‘the Carboniferous rainforest collapse’, lycopsid dominance of (mostly peat) swamps changed to marattialean tree-fern and medullosan pteridosperm dominance, and biodiversity decreased. Event 3 encompassed one glacial–interglacial cycle and included vegetational turnover in other wetland habitats. For several subsequent glacial–interglacial cycles peatland dominance varied, known from palynology, before stabilizing. These vegetational changes likely reflect climatic events driving unidirectional, non-reversible wetland vegetational changes, during cooler, wetter parts of glacial–interglacial cycles. Discussion is complicated by different placements of crucial stratigraphic boundaries, but under the same names, compromising both clear communication and understanding of the literature. Not the least is the floating base of the Cantabrian Substage, together with the position of the Westphalian–Stephanian Stage boundary.

Abstract: Forty Permian conodont biozones are summarized and 35 additional regional zones are correlated. The Lower Permian is largely zoned on the basis of partial range lineage interval zones of species of Streptognathodus , Sweetognathus , Neostreptognathodus and Mesogondolella . The Middle and Upper Permian are zoned, respectively, on the basis of partial range lineage interval zones of species of Jinogondolella and Clarkina . The ranges of all key taxa to develop this zonation are depicted in relation to the geochronological ages of Permian stages. The lowest Permian succession has been astronomically tuned with 400 kyr cyclothems linked to interpolated ages in the Uralian succession.

Abstract: The Permian Period is one of the most varied times, not only in oceanic environments, but also for the evolution of the radiolarian assemblages. Permian radiolarians have been extensively studied, and great progress has been achieved recently in their biostratigraphy. Seventeen Permian radiolarian zones are summarized here. Among these, seven zones are assigned to the latest Carboniferous–early Permian, namely (in ascending order): the Pseudoalbaillella bulbosa Assemblage Zone, the Pseudoalbaillella u-forma – Pseudoalbaillella elegans Assemblage Zone, the Pseudoalbaillella lomentaria – Pseudoalbaillella sakmarensis Assemblage Zone, the Pseudoalbaillella rhombothoracata Interval Zone, the Albaillella xiaodongensis Assemblage Zone, the Albaillella sinuata Abundance Zone and the Pseudoalbaillella ishigai Abundance Zone. Five zones are attributed to the middle Permian, including the Pseudoalbaillella globosa Interval Zone, the Follicucullus monacanthus Interval Zone, the Follicucullus porrectus Interval Zone, the Follicucullus scholasticus Interval Zone and the Follicucullus charveti Interval Zone. Five zones belong to the middle Permian–late Permian, including the Albaillella cavitata Interval Zone, the Albaillella levis Interval Zone, the Albaillella excelsa Interval Zone, the Albaillella triangularis Interval Zone and the Albaillella yaoi Abundance Zone. These Permian radiolarian biozones and their correlations with conondont zones or other chronostratigraphic schemes are discussed. The phylogenetic model of radiolarians through the Permian Period is also revised.

Abstract: Permian rugose corals underwent evolutionary episodes of assemblage changeover, biogeographical separation and extinction, which are closely related to geological events during this time. Two coral realms were recognized, the Tethyan Realm and the Cordilleran–Arctic–Uralian Realm. These are characterized by the families Kepingophyllidae and Waagenophyllidae during the Cisuralian, Waagenophyllidae in the Guadalupian and the subfamily Waagenophyllinae in the Lopingian, and the families Durhaminidae and Kleopatrinidae during the Cisuralian and major disappearance of colonial and dissepimented solitary rugose corals from the Guadalupian to the Lopingian, respectively. The development of these coral realms is controlled by the geographical barrier resulting from the Pangaea formation. According to the changes in the composition and diversity of the Permian rugose corals, a changeover event might have occurred at the end-Sakmarian and is characterized by the mixed Pennsylvanian and Permian faunas to typical Permian faunas, probably related to a global regression. In addition, three extinction events are present at the end-Kungurian, the end-Guadalupian and the end-Permian, which are respectively triggered by the northward movement of Pangaea, the Emeishan volcanic eruptions and subsequent global regression, and the global climate warming induced by the Siberian Traps eruption.