Abstract Quantitative analysis performed on latest Pliensbachian–early Toarcian calcareous nannofossil assemblages from the Camino section (Basque Cantabrian Basin) allowed their response to the environmental changes recorded during this time interval to be deciphered, characterized by an extinction event. The results were introduced within a principal component analysis and compared with the stable isotope and total organic carbon curves. During the latest Pliensbachian, the Mirabile and the lowermost part of the Semicelatum Ammonite Subzones, Schizosphaerella , Bussonius prinsii , Biscutum finchii , Calcivascularis jansae and Similiscutum avitum , taxa that probably thrived in rather cold waters, dominated the calcareous nannofossil assemblages. Coinciding with warmer and wetter conditions, which probably led to an increase in surface water fertility, recorded slightly below the extinction boundary, the mesotrophic taxa B. novum , L. hauffii and Calyculus spp. were dominant. Nevertheless, T. patulus and C. jansae , which became extinct just below the extinction boundary, show preferences for oligotrophic conditions. Salinities similar to those of modern oceans have been inferred around the extinction boundary, considering the coupling between the abundances of Calyculus spp. and the species richness together with the absence of black shales. After the extinction boundary, nannofossil assemblages were dominated by the deep-dwelling C. crassus and the shallow-dwelling Lotharingius species, interpreted as opportunistic taxa. This work confirms that calcareous nannofossils are a really useful tool for palaeoceanographic and palaeoenvironmental reconstructions, especially in terms of climatic changes.

Abstract The leading hypothesis for the Toarcian oceanic anoxic event (T-OAE; c. 183 Ma) and the associated negative C-isotope excursion is the massive release of 12 C favouring greenhouse conditions and continental weathering. The nutrient delivery to shallow basins supported productivity and, because of O 2 consumption by organic-matter respiration, anoxia development. However, several studies have shown that calcareous nannoplankton experienced a decrease during the T-OAE. Nannofossil fluxes measured in the Llanbedr (Mochras Farm) borehole, Wales, UK, were the highest prior to the negative C-isotope excursion, along with high amounts of taxa indicative of nutrient-rich environments (Biscutaceae). Such conditions attest to high productivity. Fluxes show the lowest values in the core of the event, along with a size decrease of Schizosphaerella and a peak in Calyculaceae. The recovery of nannofossil fluxes and Schizosphaerella size occurred concomitant with the return of C-isotopes to more positive values. Concomitantly, deep dwellers ( Crepidolithus crassus ) dominated, indicating a recovery of the photic-zone productivity. These observations demonstrate that the cascade of environmental responses to the initial perturbation was more complex than previously considered. In spite of elevated nutrient delivery to epicontinental basins in the early Toarcian, carbonate and primary productions of nannoplankton were depressed in the core the T-OAE, probably because of prolonged thermohaline seawater stratification.

Abstract Ophiuroids, the slender-armed cousins of starfish, constitute an important component of modern marine benthos and have been used successfully in the exploration of (palaeo)-ecological and evolutionary trends, yet their fossil record is still poorly known. One of the major gaps in the known palaeobiodiversity of this group coincides with a global palaeoenvironmental crisis during the early Toarcian (Early Jurassic, 183 myr ago), known as the Jenkyns Event. Here we describe ophiuroid remains retrieved from a series of samples from the Dudelange (Luxembourg) drill core, which spans the lower part of the Toarcian, between the top of the Pliensbachian and the onset of the Jenkyns Event. A total of 21 species are recorded, including three new genera and 12 new species. Ophiuroid diversity and abundance fluctuate in parallel with depositional facies, with lowest values coinciding with black shales. Highest diversities, including exceptional occurrences of taxa nowadays restricted to deep-sea areas, are recorded from just below the black shales, corresponding to the onset of the Jenkyns Event. Our results show that even small (100 g) bulk sediment samples retrieved from drill cores can yield numerous identifiable ophiuroid remains, thus unlocking this group for the study of faunal change across palaeoenvironmental crises.

Abstract We report new ichthyosaur material excavated in lower Toarcian levels of the LafargeHolcim Val d'Azergues quarry in Beaujolais, SE France. A partially articulated skull and a smaller, unprepared but likely subcomplete skeleton preserved in a carbonate concretion are identified as stenopterygiids, a family of wide European distribution during the Early Jurassic. These specimens are among the finest preserved Toarcian exemplars known from Europe and, in one of them, soft tissue preservation is suspected. Their state of preservation is attributed to the combination of prolonged anoxic conditions near the water–sediment interface and early carbonate cementation resulting from the activity of sulfate-reducing bacteria. We also present carbon and strontium isotope values obtained from the study site that allow detailed temporal comparisons with other Toarcian vertebrate-yielding sites and environmental perturbations associated with the Toarcian Oceanic Anoxic Event (T-OAE). These comparisons suggest that the relatively high abundance and good preservation state of Toarcian vertebrates was favoured by a prolonged period of low bottom water oxygenation and accumulation rates. The environmental conditions that prevailed during the T-OAE were probably responsible for the extensive nature of Lagerstätte-type deposits with exceptional preservation of marine organisms. Testing whether the T-OAE had a biological impact on marine vertebrates requires a precise chemostratigraphic context of the fossil record spanning the Pliensbachian–Toarcian interval.

Abstract The flooding of the Lower Jurassic shelf in the North Gondwana Palaeomargin during the early Toarcian occurred on a fragmented and irregular topography affected by differential subsidence – owing to the activity of listric faults along the North–South Axis of Tunisia – that favoured lateral changes in facies and thickness at a kilometric scale. The onset of Toarcian sedimentation (Polymorphum ammonite Zone, NJT5c nannofossil Subzone) in two adjacent sections was characterized by the deposition of limestones under high-energy conditions. The Châabet El Attaris section was located in a depressed sub-basin, and recorded restricted environmental conditions owing to water stagnation and an oxygen-depleted sea bottom. Therefore, dark mudstones developed, with increased total organic carbon contents and enhanced accumulation of redox-sensitive elements. The sedimentation of limestones bearing gutter cast structures is related to gravity flows probably linked to storm activities. These processes favoured the remobilization of sediments at the seafloor, as well as oxygen input to bottom waters, as shown by the record of trace fossils including Zoophycos , Ophiomorpha , and secondarily, Chondrites and Diplocraterion . The thinly interbedded dark mudstones are locally rich in thin-shelled bivalves that re-colonized the sea bottom after the sedimentation of these high-energy deposits, and subsequently underwent mass mortality related to the return of oxygen-depleted conditions. The Kef El Hassine section is located in the upper part of a tilted, less subsident block, as indicated by its reduced thickness compared with the Châabet El Attaris section; the absence of dark mudstones implies oxic conditions. The Polymorphum Zone consists of limestones showing evidence of sedimentation under high-energy conditions, along with hardgrounds. The occurrence of Zoophycos (deep-tiers) in the upper part of some limestone beds of the Polymorphum Zone is linked to minor erosive processes. The top of the high-energy sequence – below the deposits of a marly interval corresponding to the Levisoni Zone – is interpreted as a hardground given the high content of belemnites and Arenicolites , some of them boring on the eroded Zoophycos and Thalassinoides . This study shows that the sedimentary expression of the Jenkyns Event is not uniform across Tunisia, supporting the importance of local conditions in determining the development of anoxic conditions.

Abstract This study focuses on a condensed sequence of alternating carbonate–clastic sediments of the Barrington Member, Beacon Limestone Formation (latest Pliensbachian to early Toarcian) from Somerset (SW England). Abundant ammonites confirm (apart from the absence of the Clevelandicum and Tenuicostatum ammonite subchronozones) the presence of Hawskerense Subchronozone to Fallaciosum–Bingmanni subchronozones. Well-preserved, sometimes diverse assemblages of ostracods, foraminifera, nannofossils and low-diversity dinoflagellate assemblages support the chronostratigraphic framework. Stable-isotope analyses demonstrate the presence of a carbon isotope excursion, relating to the Toarcian Oceanic Anoxic Event, within the early Toarcian. Faunal, geochemical and sedimentological evidence suggest that deposition largely took place in a relatively deep-water (subwave base), mid-outer shelf environment under a well-mixed water column. However, reduced benthic diversity, the presence of weakly laminated sediments and changes in microplankton assemblage composition within the Toarcian Oceanic Anoxic Event indicates dysoxic, but probably never anoxic, bottom-water conditions during this event. The onset of the carbon isotope excursion coincides with extinction in the nannofossils and benthos, including the disappearance of the ostracod suborder Metacopina. Faunal evidence indicates connectivity with the Mediterranean region, not previously recorded for the UK during the early Toarcian.

Abstract In this paper, we present a detailed review of upper Pliensbachian–lower Toarcian kerogen assemblages from the southern areas of the West Tethys shelf (between Morocco and northern Spain) and demonstrate the use of the Phytoclast Group as a tracer of palaeoenvironmental changes in the early Toarcian. The kerogen assemblages in the studied sections from the southern areas of the West Tethys shelf are dominated by the Phytoclast Group and terrestrial palynomorphs, although punctual increases in amorphous organic matter, freshwater ( Botryococcus ) and marine microplankton (dinoflagellate cysts, acritarchs and prasinophyte algae) were observed at specific stratigraphic intervals. The opaque/non-opaque phytoclasts ratio was used to trace changes in palaeoclimate and other palaeoenvironmental parameters and reflect climate gradients associated with water availability during early Toarcian. During the Pliensbachian–Toarcian and Jenkyns events, changes in kerogen assemblages in the southern areas of the West Tethys shelf correlated with changes in the northern Tethys and Panthalassa shelf. The acceleration of the hydrological cycle associated with the aforementioned events was less intense in the northern Gondwana, southern and western Iberian basins, a reflection of the palaeogeographic position of these basins within the semi-arid climate belt when compared with the northern Iberian region and other northern areas of the West Tethys and Panthalassa shelf, inserted in winter-wet and warm temperate climate belts. Amorphous organic matter enrichment associated with the Pliensbachian–Toarcian and Jenkyns events reflects an increase in primary productivity linked with increased continental weathering, fluvial runoff and riverine organic matter, and nutrient input into marine areas, inducing water column stratification and promoting the preservation of organic matter.

Abstract The Toarcian Oceanic Anoxic Event (T-OAE) is marked by major palaeoenvironmental and palaeoceanographical changes on a global scale, associated with a severe disturbance of the global carbon cycle and organic-rich facies deposition. Here, a multiproxy approach (petrographic and geochemical techniques) was applied to the study of the organic content of the T-OAE of the Paris Basin, whose phytoplanktonic origin has been previously inferred by its geochemical signature. The top of the tenuicostatum Zone is characterized by palynomorphs and marine phytoplankton-derived amorphous organic matter (AOM), representing a proximal marine environment with emplacement of euxinic conditions at the top (total organic carbon/sulfur content and increase in AOM). At the base of the serpentinum Zone the proliferation of bacterial biomass begins, with phytoplankton playing a secondary role. This indicates the development of stagnant and restrictive conditions in a proximal environment, with water column stratification (neohop-13(18)-ene). The majority of the serpentinum Zone is dominated by bacterial biomass, suggesting a marine environment with bottom-water stagnation, possibly related to basin palaeogeomorphology and circulation patterns, with episodic euxinia. This therefore suggests that the T-OAE organic fraction is dominated by bacterial biomass, not phytoplankton, showing the importance of an integrated approach to the determination of the organic facies.

Abstract The historical view of an equable Jurassic greenhouse world has been challenged by recent studies documenting recurrent alternation between contrasting climate modes. Cooling of high-latitudinal areas may have been caused by orogenic processes at the northern margin of the Tethys Ocean that reduced heat transport towards the polar regions. Warm phases correlate to periods of intensified volcanism. The Jenkyns Event occurred during the transition from a late Pliensbachian icehouse into an early Toarcian greenhouse. Parallel evolution of different environmental processes, including sea level, climate and carbon cycle, indicate a causal mechanism tied to astronomical forcing. Insolation-controlled variations in the extent of the cryosphere (ice caps and permafrost) facilitated orbitally paced sea-level cycles via waxing and waning of the polar ice caps, and negative carbon isotope excursions via the release of cryosphere-bound 12 C-enriched carbon. This review and synthesis of sedimentological, geochemical and palaeontological palaeoenvironment indicators, and of simulations from climate models, aims to reconstruction, in particular, the high-latitudinal environmental conditions of late Pliensbachian–early Toarcian times. Focus is laid on the extent of the regions that were potentially suitable for hosting a cryosphere. An environmental response to cryosphere dynamics is considered to have been a key component of the Jenkyns Event.

Abstract Detailed assessment of high-resolution elemental and isotopic geochemical datasets collected from the marl–limestone alternations cropping out at La Cerradura (Subbetic domain of the Betic Cordillera, Spain) and chrono- and chemostratigraphic correlation with the reference Mochras borehole (Cardigan Bay Basin, UK) unveiled valuable new insights into understanding of late Pliensbachian–early Toarcian palaeoenvironmental dynamics in a key geographical area between the northern European seaway and the Tethys Ocean. This study shows that deposition in the study area took place under dominantly oxic water column conditions, indicated, for example, by the generalized lack of enrichment in organic matter and redox metals typically associated with anoxia and euxinia. Carbon isotope stratigraphy (δ 13 C TOC ) allowed recognition of the spinatum (= emaciatum in the Submediterranean Province), Pliensbachian–Toarcian, and early Toarcian Oceanic Anoxic Event negative carbon isotopic excursions and the late Pliensbachian positive carbon isotopic excursion. It is here suggested that the observed periodic changes in lithology and sedimentary geochemistry occur at orbital frequencies (i.e. long and short eccentricity and, tentatively, precession), hinting at an astronomical control of the local–regional climate and environment during the Pliensbachian and Toarcian in the mid- to low-latitude South Iberian palaeomargin area.

Abstract A detailed 3D petroleum system model was constructed for the Schleswig-Holstein area in northern Germany. Salt movement and the Quaternary ice episodes were implemented in order to reconstruct their impact on temperature, maturity and pressure. Burial, temperature and maturity histories were calculated for the Jurassic troughs and the Glueckstadt Graben showing both differences and similarities. For example, all locations reached (almost) deepest burial at present day, whilst subsidence and long-term sedimentation rate was highest in Glueckstadt Graben during the Triassic. The Jurassic troughs received their major subsidence and sedimentation pulse later, and were strongly affected by a later salt movement. The implementation of Quaternary glacial episodes does not have a strong impact on petroleum generation from the major source rock (Lower Toarcian Posidonia Shale). In the case of the Posidonia Shale reaching the stage of petroleum expulsion (outside of the study area), the effect of ‘glacial pumping’(i.e. the development of high pore pressures during glaciation followed by expulsion and subsequent pressure release during deglaciation) can be deduced from the model. Petroleum accumulations in the reservoir layers (Dogger sandstones) are also seen to have been affected. This finding is of interest for exploration, as it might control petroleum composition, biodegradation and leakage through cap rocks.