ABSTRACT The Franciscan subduction complex formed over a protracted, ~150 m.y. period, during Late Jurassic to late Cenozoic subduction of oceanic lithosphere beneath the western margin of the North American continent. Growth of the complex occurred chiefly by progressive accretion, in which voluminous sediment was eroded from the magmatic arc and continent, deposited in the trench region, and then progressively subducted and accreted soon after deposition. The Yolla Bolly terrane, a major Franciscan subunit, has stood out as a possible exception to a progressive-accretion model. Yolla Bolly clastic rocks are almost barren of fossils, but there are ~13 localities with Late Jurassic and Early Cretaceous Buchia specimens, ~3 with mid-Cretaceous ammonites or Inoceramus , plus several with mid-Cretaceous youngest detrital-zircon populations. These ages had suggested that sediments may not have been deposited into an active trench, but instead were deposited into a relatively stable Yolla Bolly basin, which was both long-lived (ca. 150 Ma through ca. 95 Ma) and far-traveled (exotic). This basin was then accreted and metamorphosed at perhaps 92 Ma. It is surprising, however, that such a basin could have survived for ~50 m.y. along a subduction margin before being accreted. We determined detrital-zircon U-Pb ages from 31 new sandstone samples, including from key Buchia sites, and they indicate that Yolla Bolly clastic deposition actually occurred almost entirely between ca. 115 and 98 Ma. All of the Buchia specimens in the main parts of the Yolla Bolly terrane have been redeposited and the arc- and continent-sourced clastic rocks that comprise almost all of the terrane are much younger than once thought. This makes evolution of the Yolla Bolly terrane compatible with a progressive-accretion model, in which its constituent packets of clastic rocks were deposited in a native trench setting and then rapidly subducted, accreted, and metamorphosed.

One of the rarest of marine reptiles is the mosasaur genus Globidens , characterized by a massive, bulbous dentition. The rarity of the taxon, coupled with the bulbous dentition, resulted in various theories concerning life habits. Although a consensus indicates that the dentition was adapted for crushing resistant elements, hypotheses have varied concerning prey, ranging from turtles or bivalves to scavenging. Finally, a partial skeleton of Globidens has been recovered from the Big Bend area of the Missouri River in central South Dakota. The specimen was discovered in the upperDeGrey Formation (upperCampanian) of the Pierre Shale Group. During analysis, bivalve fragments were found packed within the rib-cage region of the skeleton. In the field, bivalve concentrations did not occur laterally or above or below the skeleton, indicating that they were the stomach contents of the mosasaur. Associated within the stomach area are a number of bivalve taxa, including oysters and small bivalves with lamellar shells, probably of the genus Anomia . The most common specimens within the stomach area are bivalves that exhibit a prismatic shell microstructure typical of inoceramids. Four inoceramid shell morphotypes were recovered, including a coarse-ribbed morphotype, a fine-ribbed morphotype, one with a thickened umbo, and a large, flat, thin-shelled morphotype. Because of their position in the mosasaur, their fragmented condition, limited taxonomic diversity, and absence from surrounding sediments, the bivalves are considered stomach contents. Some smaller, complete shells of Anomia escaped breakage, whereas larger inoceramids were invariably crushed. Chondrichthyan teeth were found associated but are interpreted to be the result of scavenging. This specimen of Globidens appears to have had a preference forthe large, flat, relatively thin-shelled inoceramids that contained a large, fleshy visceral mass.