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Pygidia of Lonchopygella megaspina Zhou, 1977. 1, meraspid-H4, NIGPAS151682, ×22; 2, meraspid-H3, NIGPAS151683, ×20; 3, meraspid-H3, NIGPAS151684, ×16; 4, meraspid-H3, NIGPAS151702, ×18; 5, meraspid-H2, NIGPAS151685, ×21; 6, meraspid-H2, NIGPAS151686, ×14.6; 7, meraspid-H2, NIGPAS151703, ×15; 8, meraspid-H2, NIGPAS151704, ×12.4; 9, meraspid-H1, NIGPAS151687, ×11; 10, early holaspis, NIGPAS151705, ×12; 11, early holaspis, NIGPAS151706, ×7.6; 12, early holaspis, NIGPAS151707, ×12; 13, later holaspis, NIGPAS151708, ×5; 14, later holaspis, NIGPAS151709, ×10.4; 15, later holaspis, NIGPAS151688, ×11.4; 16, later holaspis, NIGPAS151689, ×7.8; 17, later holaspis, NIGPAS151690, ×3.5; 18, later holaspis, NIGPAS151691, ×7.5; 19, later holaspis, NIGPAS151692, ×7.6; 20, later holaspis, NIGPAS151693, ×7.2; 21, later holaspis, NIGPAS151694, ×7.5; 22, later holaspis, NIGPAS151695, ×8.3; 23, later holaspis, NIGPAS151696, ×9; 24, later holaspis, NIGPAS151697, ×5.7; 25, later holaspis, NIGPAS151698, ×6.9; 26, later holaspis, NIGPAS151711, ×5.4; 27, 28, later holaspis, NIGPAS151701, dorsal and left lateral view respectively, ×3.3; 29, later holaspis, NIGPAS151710, ×6; 30, later holaspis, NIGPAS151712, ×6.3; 31, later holaspis, NIGPAS151699, ×3.9; 32, later holaspis, NIGPAS151700, ×5.2.
Published: 01 May 2013
-H2, NIGPAS151703, ×15; 8 , meraspid-H2, NIGPAS151704, ×12.4; 9 , meraspid-H1, NIGPAS151687, ×11; 10 , early holaspis, NIGPAS151705, ×12; 11 , early holaspis, NIGPAS151706, ×7.6; 12 , early holaspis, NIGPAS151707, ×12; 13 , later holaspis, NIGPAS151708, ×5; 14 , later holaspis, NIGPAS151709
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Otarionella rastrum sp. nov. (a–m, t–v) Scanning electron micrographs of silicified specimens from the middle member of the Bissounel Formation (early to late Emsian), Bissounel Peak, Montagne Noire, France. (n–s, w–y) Digital photographs of calcareous specimens from the Mont Peyroux Formation (late Emsian), Montagne Noire, France. Scale bars 0.25 mm (a–e) and 0.5 mm (f–y). (a–e) Metaprotaspides; (a) dorsal view, UMC-IP490; (b) dorsal view, UMC-IP491; (c–e) specimen with anterior border broken, UMC-IP492; (c) dorsal view; (d) left lateral view; (e) postero-lateral view. (f–r) Cranidia, dorsal views; (f) smallest meraspid specimen, UMC-IP493; (g) meraspis, UMC-IP494; (h) meraspis, UMC-IP495; (i) meraspis, UMC-IP496; (j) meraspis, UMC-IP497; (k) meraspis (?), UMC-IP498; (l) meraspis (?), UMC-IP499; (m) meraspis (?), UMC-IP500; (n) holaspis (?), UMC-IP501; (o) holaspis, UMC-IP502; (p) holaspis, UMC-IP503; (q) holotype, holaspis, UMC-IP504; (r) holaspis, UMC-IP505; (s) holaspis, anterior view, UMC-IP506. (t, u) Fragments of the anterior region, anterior views; (t) small specimen, UMC-IP507; (u) large specimen, UMC-IP508. (v–x) Right lateral views; (v) meraspis, UMC-IP496; (w) holaspis (?), UMC-501; (x) holaspis, UMC-IP502; (y) mirrored left lateral view, holaspis, UMC-IP505.
Published: 01 January 2008
, dorsal views; (f) smallest meraspid specimen, UMC-IP493; (g) meraspis, UMC-IP494; (h) meraspis, UMC-IP495; (i) meraspis, UMC-IP496; (j) meraspis, UMC-IP497; (k) meraspis (?), UMC-IP498; (l) meraspis (?), UMC-IP499; (m) meraspis (?), UMC-IP500; (n) holaspis (?), UMC-IP501; (o) holaspis, UMC-IP502; (p
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Drevermannia antecurvata sp. nov., upper Famennian (to V), lower part of the Breternitz Member, Breternitz locality, Thuringia, Germany. All figures are scanning electron micrographs of silicified specimens. Scale bars 0.5 mm. (a–h) Cranidia, dorsal views. (a) Smallest meraspis, MB.T.4867.1. (b) Meraspis, MB.T.4867.2. (c) Meraspis, MB.T.4867.3. (d) Meraspis, MB.T.4867.4. (e) Meraspis, MB.T.4867.5. (f) Holaspis (?), MB.T.4867.6. (g) Large holaspis, MB.T.4867.7. (h) Holotype, large holaspis, MB.T.4867.8. (i) Right librigena, ventral view, MB.T.4867.9. (j–n) Librigenae, dorsal views. (j) Smallest specimen, MB.T.4867.10. (k) MB.T.4867.11. (l) MB.T.4867.12. (m) MB.T.4867.13. (n) Largest specimen, MB.T.4867.14. (o, p) Hypostomes, ventral views. (o) Small specimen, MB.T.4867.15. (p) Large specimen, MB.T.4867.16. (q–w) Transitory pygidia and pygidia, dorsal views. (q) Smallest meraspis, MB.T.4867.17. (r) Meraspis, MB.T.4867.18. (s) Meraspis, MB.T.4867.19. (t) Meraspis (?), MB.T.4867.20. (u) Holaspis, MB.T.4867.21. (v) Holaspis (right side broken), MB.T.4867.22. (w) Largest holaspis (left anterior part broken), MB.T.4867.23. (x, y) Transitory pygidium and pygidium, ventral views. (x) Meraspis, MB.T.4867.24. (y) Holaspis, MB.T.4867.25.
Published: 01 January 2006
.4867.1. (b) Meraspis, MB.T.4867.2. (c) Meraspis, MB.T.4867.3. (d) Meraspis, MB.T.4867.4. (e) Meraspis, MB.T.4867.5. (f) Holaspis (?), MB.T.4867.6. (g) Large holaspis, MB.T.4867.7. (h) Holotype, large holaspis, MB.T.4867.8. (i) Right librigena, ventral view, MB.T.4867.9. (j–n) Librigenae, dorsal views. (j
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Figure 4. Small (A) to large (L) specimens of Triarthrus becki. A, Meraspis, NYSM 16781, Cephalic length (CL) = 0.97 mm. B, Meraspis, NYSM 16782, CL = 1.13 mm. C, Meraspis, NYSM 16783, CL = 1.31 mm. D, Meraspis, NYSM 16784, CL = 1.41 mm. E, Meraspis, NYSM 16785, CL = 1.49 mm. F, Meraspis, NYSM 17058, CL = 1.72 mm. G, Meraspis, NYSM 17059, CL = 1.91 mm. H, Meraspis, NYSM 17060, CL = 2.13 mm. I, Holaspis, NYSM 17061, CL = 3.10 mm. J, Holaspis, NYSM 17062, CL = 4.25 mm. K, Holaspis, NYSM 17063, CL = 5.39 mm. L, Holaspis, NYSM 17064, CL = 6.42 mm. These 12 specimens are reposited in the New York State Museum as hypotypes
Published: 01 January 2002
17058, CL = 1.72 mm. G, Meraspis, NYSM 17059, CL = 1.91 mm. H, Meraspis, NYSM 17060, CL = 2.13 mm. I, Holaspis, NYSM 17061, CL = 3.10 mm. J, Holaspis, NYSM 17062, CL = 4.25 mm. K, Holaspis, NYSM 17063, CL = 5.39 mm. L, Holaspis, NYSM 17064, CL = 6.42 mm. These 12 specimens are reposited in the New York
Journal Article
Journal: Paleobiology
Published: 01 January 2002
Paleobiology (2002) 28 (3): 364–377.
... 17058, CL = 1.72 mm. G, Meraspis, NYSM 17059, CL = 1.91 mm. H, Meraspis, NYSM 17060, CL = 2.13 mm. I, Holaspis, NYSM 17061, CL = 3.10 mm. J, Holaspis, NYSM 17062, CL = 4.25 mm. K, Holaspis, NYSM 17063, CL = 5.39 mm. L, Holaspis, NYSM 17064, CL = 6.42 mm. These 12 specimens are reposited in the New York...
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Otarionella rastrum sp. nov. (a–g, j–y, aa, bb) Scanning electron micrographs of silicified specimens from the middle member of the Bissounel Formation (early to late Emsian), Bissounel Peak, Montagne Noire, France. (h, i, z, cc) Digital photographs of calcareous specimens from the Mont Peyroux Formation (late Emsian), Montagne Noire, France. Scale bars 0.5 mm (a–s) and 0.25 mm (t–cc). (a–g) Librigenae, dorsal views; (a) smallest meraspid specimen, UMC-IP509; (b) meraspis, UMC-IP510; (c) meraspis (mirrored), UMC-IP511; (d) meraspis, UMC-IP512; (e) meraspis (?), UMC-IP513; (f) meraspis (?) (mirrored), UMC-IP514; (g) holaspis (?) (mirrored), UMC-IP515. (h, i) Librigenae, dorso-lateral views; (h) holaspis with genal spine broken, UMC-IP516; (i) holaspis with lateral border broken (mirrored), UMC-IP517. (j) Right librigena, ventral view (mirrored), UMC-IP518. (k) Left librigena (note the three rows of lateral border spines), lateral view, UMC-IP519. (l–n) Hypostomes, ventral views; (l) UMC-IP520; (m) UMC-IP521; (n) UMC-IP522. (o–s) Thoracic segments with (o, q, s) or without (p, r) an axial spine; (o) right lateral view, UMC-IP523; (p) posterior view, UMC-IP524; (q) anterior view, UMC-IP525; (r) dorsal view (front to the top), UMC-IP526; (s) dorsal view (front to the top), UMC-IP527. (t–z) Transitory pygidia and pygidia, dorsal views; (t) meraspid degree 2, UMC-IP528; (u) meraspid degree 3, UMC-IP529; (v) large meraspid degree 3, UMC-IP530; (w) meraspid degree 4, UMC-IP531; (x) meraspis, UMC-IP532; (y) small holaspis (?), UMC-IP533; (z) large holaspis, UMC-IP534. (aa) Small meraspis, ventral view, UMC-IP535; (bb) meraspid degree 2, lateral view (mirrored), UMC-IP528; (cc) large holaspis, lateral view, UMC-IP534.
Published: 01 January 2008
) meraspis (?) (mirrored), UMC-IP514; (g) holaspis (?) (mirrored), UMC-IP515. (h, i) Librigenae, dorso-lateral views; (h) holaspis with genal spine broken, UMC-IP516; (i) holaspis with lateral border broken (mirrored), UMC-IP517. (j) Right librigena, ventral view (mirrored), UMC-IP518. (k) Left librigena
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Figure5—Telopeltis microphthalmus n. sp., Virgin Hills Formation, Late Frasnian Zone 13, from McWhae Ridge, Lawford Range, Canning Basin, Western Australia. 1, Cranidium, early holaspis, WAM 04.305, dorsal view, ×8.5; 2–4, cranidium, WAM 04.306, dorsal, lateral, and frontal views, ×6.6; 5, librigena, WAM 96.474, dorsal view, ×5.3; 6–8, holotype, cephalon, WAM 96.479, dorsal, frontal, and lateral views, ×6.4; 9, 10, pygidium, WAM 04.307, lateral and dorsal views, ×5.7; 11, cranidium, early holaspis, WAM 04.308, dorsal view, ×8.3; 12–14, fragmentary pygidium, WAM 04.309, frontal, lateral, and dorsal views, ×4.9; 15, 16, pygidium, early holaspis, WAM 04.310, dorsal and lateral views, ×7.
Published: 01 September 2006
Figure 5 — Telopeltis microphthalmus n. sp., Virgin Hills Formation, Late Frasnian Zone 13, from McWhae Ridge, Lawford Range, Canning Basin, Western Australia. 1, Cranidium, early holaspis, WAM 04.305, dorsal view, ×8.5; 2–4, cranidium, WAM 04.306, dorsal, lateral, and frontal views, ×6.6; 5
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Drevermannia sp. 1, upper Famennian (to V), lower part of the Breternitz Member, Breternitz locality, Thuringia, Germany. All figures are scanning electron micrographs of silicified specimens. Scale bars 0.5 mm. (a) Metaprotaspis, dorsal view, MB.T.4868.1. (b, c, e–g) Cranidia, dorsal views. (b) Small meraspis, MB.T.4868.2. (c) Meraspis, MB.T.4868.3. (e) Meraspis, MB.T.4868.4. (f) Holaspis (?), MB.T.4868.5. (g) Holaspis (?), MB.T.4868.6. (d, h) Librigenae, dorsal views. (d) Small specimen, MB.T.4868.7. (h) Large specimen, MB.T.4868.8.
Published: 01 January 2006
, dorsal views. (b) Small meraspis, MB.T.4868.2. (c) Meraspis, MB.T.4868.3. (e) Meraspis, MB.T.4868.4. (f) Holaspis (?), MB.T.4868.5. (g) Holaspis (?), MB.T.4868.6. (d, h) Librigenae, dorsal views. (d) Small specimen, MB.T.4868.7. (h) Large specimen, MB.T.4868.8.
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Figure3—Telopeltis woodwardi n. sp., Virgin Hills Formation, Middle Frasnian Zone 12, from McWhae Ridge, Lawford Range, Canning Basin, Western Australia. 1–3, Cranidium, WAM 91.306, dorsal, lateral, and frontal views, ×5; 4–6, cranidium, early holaspis, WAM 04.298, dorsal, frontal, and lateral views, ×7.9; 7, eye lobe, WAM 04.299, dorsal view, ×5.8; 8, fragmentary cranidium, internal mold, WAM 04.300, dorsal view, ×3.5; 9–12, pygidium, WAM 04.301, frontal, dorsal, posterior, and lateral views, ×3; 13, fragmentary pygidium, WAM 04.302, dorsal view, ×3.3; 14, fragmentary pygidium, late holaspis, WAM 04.303, lateral view, ×2.7; 15–18, holotype, pygidium, WAM 04.304, frontal, dorsal, posterior, and lateral views, ×3.
Published: 01 September 2006
Figure 3 — Telopeltis woodwardi n. sp., Virgin Hills Formation, Middle Frasnian Zone 12, from McWhae Ridge, Lawford Range, Canning Basin, Western Australia. 1–3, Cranidium, WAM 91.306, dorsal, lateral, and frontal views, ×5; 4–6, cranidium, early holaspis, WAM 04.298, dorsal, frontal
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Drevermannia richteriPfeiffer, 1959, upper Famennian (to VI), upper part of the Breternitz Member, Breternitz and Fischersdorf localities, Thuringia, Germany. All figures are scanning electron micrographs of silicified specimens. Scale bars 0.5 mm. (a–f) Librigenae, dorsal views. (a) Smallest specimen, MB.T.4866.22. (b) MB.T.4866.23. (c) MB.T.4866.24. (d) MB.T.4866.25. (e) MB.T.4866.26. (f) Largest specimen, MB.T.4866.27. (g) Left librigena, ventral view, MB.T.4866.28. (h) Left librigena, lateral view, MB.T.4866.29. (i) Large pygidium, ventral view, MB.T.4866.30. (j–n) Transitory pygidia and pygidia, dorsal views. (j) Small meraspis, MB.T.4866.31. (k) Meraspis, MB.T.4866.32. (l) Meraspis (?), MB.T.4866.33. (m) Holaspis, MB.T.4866.34. (n) Largest holaspis, MB.T.4866.35.
Published: 01 January 2006
pygidia and pygidia, dorsal views. (j) Small meraspis, MB.T.4866.31. (k) Meraspis, MB.T.4866.32. (l) Meraspis (?), MB.T.4866.33. (m) Holaspis, MB.T.4866.34. (n) Largest holaspis, MB.T.4866.35.
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Pygidium segmentation schedule for Lonchopygella megaspina Zhou, 1977. Abbreviations: H=holaspis, M=meraspis, M-Hn=number of the meraspid instar prior to onset of the holaspid stage (see text for details).
Published: 01 May 2013
Figure 4 Pygidium segmentation schedule for Lonchopygella megaspina Zhou, 1977. Abbreviations: H=holaspis, M=meraspis, M-Hn=number of the meraspid instar prior to onset of the holaspid stage (see text for details).
Journal Article
Published: 01 May 2013
Journal of Paleontology (2013) 87 (3): 472–483.
...-H2, NIGPAS151703, ×15; 8 , meraspid-H2, NIGPAS151704, ×12.4; 9 , meraspid-H1, NIGPAS151687, ×11; 10 , early holaspis, NIGPAS151705, ×12; 11 , early holaspis, NIGPAS151706, ×7.6; 12 , early holaspis, NIGPAS151707, ×12; 13 , later holaspis, NIGPAS151708, ×5; 14 , later holaspis, NIGPAS151709...
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Reconstructions of various ontogenetic stages of Redlichia cf. versabunda from the Ramsay Limestone (Yorke Peninsula, South Australia): (a) early-stage protaspis; (b) late-stage protaspis; (c–g) meraspid stages in order of increasing size; and (h) holaspis. Dashed lines represent uncertain morphology.
Published: 02 December 2020
Fig. 8. Reconstructions of various ontogenetic stages of Redlichia cf. versabunda from the Ramsay Limestone (Yorke Peninsula, South Australia): (a) early-stage protaspis; (b) late-stage protaspis; (c–g) meraspid stages in order of increasing size; and (h) holaspis. Dashed lines represent
Journal Article
Journal: Paleobiology
Published: 01 November 2018
Paleobiology (2018) 44 (4): 638–659.
... gape mode, characterized by opening of the facial sutures and separation of the librigenae, they disarticulated the entire cephalon in Salter’s mode of molting. For other phacopine clades (Phacopoidea) the transition to Salter’s mode occurs during the meraspid period or at the onset of holaspis...
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Journal Article
Published: 01 January 2006
Geological Magazine (2006) 143 (1): 89–104.
....4867.1. (b) Meraspis, MB.T.4867.2. (c) Meraspis, MB.T.4867.3. (d) Meraspis, MB.T.4867.4. (e) Meraspis, MB.T.4867.5. (f) Holaspis (?), MB.T.4867.6. (g) Large holaspis, MB.T.4867.7. (h) Holotype, large holaspis, MB.T.4867.8. (i) Right librigena, ventral view, MB.T.4867.9. (j–n) Librigenae, dorsal views. (j...
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Journal Article
Published: 01 January 2008
Geological Magazine (2008) 145 (1): 55–71.
..., dorsal views; (f) smallest meraspid specimen, UMC-IP493; (g) meraspis, UMC-IP494; (h) meraspis, UMC-IP495; (i) meraspis, UMC-IP496; (j) meraspis, UMC-IP497; (k) meraspis (?), UMC-IP498; (l) meraspis (?), UMC-IP499; (m) meraspis (?), UMC-IP500; (n) holaspis (?), UMC-IP501; (o) holaspis, UMC-IP502; (p...
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Journal Article
Published: 01 September 2006
Journal of Paleontology (2006) 80 (5): 981–992.
...Figure 5 — Telopeltis microphthalmus n. sp., Virgin Hills Formation, Late Frasnian Zone 13, from McWhae Ridge, Lawford Range, Canning Basin, Western Australia. 1, Cranidium, early holaspis, WAM 04.305, dorsal view, ×8.5; 2–4, cranidium, WAM 04.306, dorsal, lateral, and frontal views, ×6.6; 5...
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Journal Article
Published: 01 May 2006
Journal of Paleontology (2006) 80 (3): 529–536.
..., the shape change between a young (probably meraspis M10) and an older specimen (holaspis) corresponds with shortening (sag.) of the cephalon, extension (tr.) of the glabella, slight elongation of the visual surface and its migration backwards and outwards, and reduction of the genal spine during ontogeny...
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All specimens are from Furongian sediments of the McKay Group, Locality 15, Wujiajiania lyndasmithae Subzone of Elvinia Zone, Bull River Valley region, southeastern British Columbia. (1, 3–5, 9) Proceratopyge rectispinata (Troedsson, 1937): (1) dorsal view of articulated molt RBCM-P954; (3) dorsal view of articulated carcass carapace RBCM-P956; (4) dorsal view of incomplete pygidium RBCM-P957; (5) dorsal view of articulated carcass carapace RBCM-P958; (9) dorsal view of articulated molt cranidium and thoracopygon, with free cheek of Olenaspella chrisnewi Chatterton and Gibb, 2016, RBCM-P959. (2) Cernuolimbus ludvigseni Chatterton and Gibb, 2016; dorsal view of carcass carapace (shell largely missing) RBCM-P955, showing a ventral median suture. (6) Proceratopyge canadensis (Chatterton and Ludvigsen, 1998); dorsal view of articulated carcass carapace RBCM-P960. (7, 8) Housia vacuna Walcott, 1890: (7) dorsal view of slightly disarticulated molt carapace RBCM-P961, X2.8 (note this specimen is coated with crystals of calcite); (8) dorsal view of articulated carcass carapace of small holaspis RBCM-P962. Scale bars = 5 mm.
Published: 01 July 2020
carapace RBCM-P960. ( 7, 8 ) Housia vacuna Walcott, 1890 : ( 7 ) dorsal view of slightly disarticulated molt carapace RBCM-P961, X2.8 (note this specimen is coated with crystals of calcite); ( 8 ) dorsal view of articulated carcass carapace of small holaspis RBCM-P962. Scale bars = 5 mm.
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All specimens are from Furongian sediments of the McKay Group, Wujiajiania lyndasmithae Subzone of Elvinia Zone, Bull River Valley region, southeastern British Columbia. All figures are of specimens from Locality 15, except (7), which is from Locality 7B of Chatterton and Gibb (2016, text-figs. 1, 4). (1, 4) Cernuolimbus ludvigseni Chatterton and Gibb, 2016: (1) dorsal view of partly articulated partial molt specimen RBCM-P963; (4) dorsal view of incomplete articulated carcass carapace RBCM-P964. (2, 5) Wujiajiania lyndasmithae Chatterton and Gibb, 2016: (2) dorsal view of articulated carcass specimen RBCM-P967; (5) ventral view of articulated carcass carapace RBCM-P968. (3, 6) Irvingella species B Chatterton and Gibb, 2016: (3) mostly dorsal view of articulated molt thoracopygon and disarticulated free cheek (in ventral view) RBCM-P965; (6) dorsal view of partly articulated molt RBCM-P966. (7, 9) Pseudeugonocare bispinatum (Kobayashi, 1962): (7) Ventral view of incomplete articulated carcass specimen RBCM-P969; (9) dorsal view of incomplete articulated molt with cranidium attached to more anterior 22 thoracic segments RBCM-P970. (8) Irvingella species; dorsal view of small holaspis or large meraspis RBCM-P971. (1–4, 6, 7, 9) Scale bars = 10 mm; (5, 8) scale bars = 2.5 mm.
Published: 01 July 2020
. ( 8 ) Irvingella species; dorsal view of small holaspis or large meraspis RBCM-P971. ( 1–4, 6, 7, 9 ) Scale bars = 10 mm; ( 5, 8 ) scale bars = 2.5 mm.