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Wilbertopora

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Journal Article

MORPHOLOGICAL DIFFERENTIATION OF AVICULARIA AND THE PROLIFERATION OF SPECIES IN MID-CRETACEOUS WILBERTOPORA (BRYOZOA: CHEILOSTOMATA) Available to Purchase

ALAN H. CHEETHAM, JOANN SANNER, PAUL D. TAYLOR, ANDREW N. OSTROVSKY
Published: 01 January 2006
Journal of Paleontology (2006) 80 (1): 49–71.
DOI: https://doi.org/10.1666/0022-3360(2006)080[0049:MDOAAT]2.0.CO;2
...ALAN H. CHEETHAM; JOANN SANNER; PAUL D. TAYLOR; ANDREW N. OSTROVSKY Abstract Discovery of avicularium-like polymorphs in Wilbertopora mutabilis Cheetham, 1954 has provided not only a new opportunity for revising the genus Wilbertopora Cheetham, 1954 , but also a more detailed basis for documenting...
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View PDF for article title, MORPHOLOGICAL DIFFERENTIATION OF AVICULARIA AND THE PROLIFERATION OF SPECIES IN MID-CRETACEOUS <span class="search-highlight">WILBERTOPORA</span> (BRYOZOA: CHEILOSTOMATA)
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 Figure3—Wilbertopora spp., Late Campanian, Point Loma Fm, Carlsbad Research Center Phase 1, San Diego County, California. 1–5,Wilbertopora sannerae sp. nov., SDNHM 26077, holotype; 1, general view of the colony, ×13; 2, ovicellate autozooids and two avicularia, that on the left with an intramural bud, ×50; 3, ovicell, ×190; 4, autozooids and avicularia; one of the autozooids (lower centre) hosts an intramural bud and is succeeded distally by a kenozooid, ×52. 5, Ovicellate autozooid flanked by two avicularia, ×72. 6,Wilbertopora sp. 1, SDNHM 26204, autozooids and small adventitious avicularia, ×60.

Figure 3 — Wilbertopora spp., Late Campanian, Point Loma Fm, Carlsbad Res... Available to Purchase

Published: 01 July 2008
Figure 3 — Wilbertopora spp., Late Campanian, Point Loma Fm, Carlsbad Research Center Phase 1, San Diego County, California. 1–5, Wilbertopora sannerae sp. nov., SDNHM 26077, holotype; 1, general view of the colony, ×13; 2, ovicellate autozooids and two avicularia, that on the left
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Table 3 —Measurements (mm) of Washita Group Wilbertopora species Available to Purchase

Published: 01 January 2006
Table 3 —Measurements (mm) of Washita Group Wilbertopora species
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Table 4 —Measurements (mm) of Washita Group Wilbertopora species Available to Purchase

Published: 01 January 2006
Table 4 —Measurements (mm) of Washita Group Wilbertopora species
Journal Article

Some Calloporid (Bryozoa: Cheilostomata) Species from the Cenozoic Sediments of Western Kachchh, Gujarat Available to Purchase

M. A. Sonar, R. V. Pawar
Published: 01 July 2016
Jour. Geol. Soc. India (2016) 88 (1): 47–54.
DOI: https://doi.org/10.1007/s12594-016-0457-y
... and Wilbertopora sp. are new and the other species viz; Crassimarginatella blandfordi and Reptoporina chhasraensis are reported for the second time from these sediments. The phylogenetic analysis suggests all the species of Calloporid bryozoans are monophyletic i.e. all of these species have common ancestor...
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Journal Article

Late Cretaceous Cheilostome Bryozoans from California and Baja California Available to Purchase

Paul D. Taylor
Published: 01 July 2008
Journal of Paleontology (2008) 82 (4): 823–834.
DOI: https://doi.org/10.1666/07-033.1
...Figure 3 — Wilbertopora spp., Late Campanian, Point Loma Fm, Carlsbad Research Center Phase 1, San Diego County, California. 1–5, Wilbertopora sannerae sp. nov., SDNHM 26077, holotype; 1, general view of the colony, ×13; 2, ovicellate autozooids and two avicularia, that on the left...
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View PDF for article title, Late Cretaceous Cheilostome Bryozoans from California and Baja California
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Table 2 —Numbers of colonies in formations of the Washita Group in northea... Available to Purchase

Published: 01 January 2006
Table 2 —Numbers of colonies in formations of the Washita Group in northeastern Texas and southeastern Oklahoma assigned to each of the eight species of Wilbertopora . Numbers comprise all of the type and other material listed in the systematic paleontology section
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 Figure4—Wilbertopora sp. 2, SDNHM 111478, Maastrichtian, Cabrillo Fm, Point Loma, San Diego County, California; 1, autozooids, some with ovicells, and a questionable avicularium (arrowed), ×44; 2, ovicellate autozooid, ×72.

Figure 4 — Wilbertopora sp. 2, SDNHM 111478, Maastrichtian, Cabrillo Fm, ... Available to Purchase

Published: 01 July 2008
Figure 4 — Wilbertopora sp. 2, SDNHM 111478, Maastrichtian, Cabrillo Fm, Point Loma, San Diego County, California; 1, autozooids, some with ovicells, and a questionable avicularium (arrowed), ×44; 2, ovicellate autozooid, ×72.
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Figure3—Plots of eight species of Wilbertopora on first three canonical axes from discriminant analysis based on 60 characters, together accounting for more than 80% of the total variance. 1, Canonical variates 1 and 2. 2, Canonical variates 1 and 3

Figure 3 —Plots of eight species of Wilbertopora on first three canonica... Available to Purchase

Published: 01 January 2006
Figure 3 —Plots of eight species of Wilbertopora on first three canonical axes from discriminant analysis based on 60 characters, together accounting for more than 80% of the total variance. 1, Canonical variates 1 and 2. 2, Canonical variates 1 and 3
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Figure4—Two hypotheses of relationship among the eight species of Wilbertopora. The trees, rooted on W. mutabilis, are maximally parsimonious, with length 122, consistency index 0.6721, and retention index 0.5604. Trees, calculated in PAUP* (Swofford, 2000), were plotted in Treeview (Page, 1996)

Figure 4 —Two hypotheses of relationship among the eight species of Wilbe... Available to Purchase

Published: 01 January 2006
Figure 4 —Two hypotheses of relationship among the eight species of Wilbertopora . The trees, rooted on W. mutabilis , are maximally parsimonious, with length 122, consistency index 0.6721, and retention index 0.5604. Trees, calculated in PAUP* ( Swofford, 2000 ), were plotted in Treeview ( Page
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Table 1 —Discrimination of eight species of Wilbertopora based on 60 met... Available to Purchase

Published: 01 January 2006
Table 1 —Discrimination of eight species of Wilbertopora based on 60 metric characters, of which 18 contributed significantly to the final DA (Appendix 2). Above diagonal , morphological distance, square root of Mahalanobis D 2 . Below diagonal , F-value for multivariate distinctness, each
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Figure5—Wilbertopora sp., USNM 216144, HTL-170, Tarrant County, Texas, Kiamichi Formation, Upper Albian. 1, Ordinary and ovicell-bearing autozooids, and three avicularia resembling those of W. tappanae n. sp. (see Fig. 9.5). 2, Another part of same colony with ovicell-bearing autozooids and avicularium resembling those of W. spatulifera n. sp. (see Fig. 10.2). Scale bar 100 μm

Figure 5 — Wilbertopora sp., USNM 216144, HTL-170, Tarrant County, Texas,... Available to Purchase

Published: 01 January 2006
Figure 5 — Wilbertopora sp., USNM 216144, HTL-170, Tarrant County, Texas, Kiamichi Formation, Upper Albian. 1, Ordinary and ovicell-bearing autozooids, and three avicularia resembling those of W. tappanae n. sp. (see Fig. 9.5 ). 2, Another part of same colony with ovicell-bearing
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Figure6—Wilbertopora listokinae n. sp., Grayson County, Texas, Grayson Formation, Lower Cenomanian. 1, Paratype, USNM 216140, HTL-5, ordinary autozooids in zone of astogenetic change, three with closure plates, and ordinary autozooids, one budded intramurally, in zone of repetition. 2, Holotype, USNM 216175, HTL-5, ovicell-bearing and maternal autozooids, and one avicularium proximal to ovicell and apparently occluding its opening. 3, Paratype, USNM 526269, HTL-5, ordinary autozooid and avicularium. 4, Paratype, NHM BZ1856(1), ordinary and ovicell-bearing autozooids with calcified closure plates. 5, Paratype, NHM BZ1627, two ordinary autozooids within which other zooids have been budded intramurally, one with a calcified closure plate. Scale bar 100 μm

Figure 6 — Wilbertopora listokinae n. sp., Grayson County, Texas, Grayson... Available to Purchase

Published: 01 January 2006
Figure 6 — Wilbertopora listokinae n. sp., Grayson County, Texas, Grayson Formation, Lower Cenomanian. 1, Paratype, USNM 216140, HTL-5, ordinary autozooids in zone of astogenetic change, three with closure plates, and ordinary autozooids, one budded intramurally, in zone of repetition. 2
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Figure7—Wilbertopora mutabilis, Fort Worth Formation, Upper Albian. 1, Holotype, LSU 4500, Denton County, Texas; ordinary autozooids, zone of astogenetic change. 2, Holotype, LSU 4500, Denton County, Texas; ordinary autozooids, avicularian polymorphs, and two ovicells, one of which is developed on a polymorph. 3, USNM 216155, HTL-67, Marshall County, Oklahoma; avicularian polymorph that apparently also served as a maternal autozooid; note partially developed (or broken) ovicell on its proximal gymnocyst. Scale bars 100 μm

Figure 7 — Wilbertopora mutabilis , Fort Worth Formation, Upper Albian. 1... Available to Purchase

Published: 01 January 2006
Figure 7 — Wilbertopora mutabilis , Fort Worth Formation, Upper Albian. 1, Holotype, LSU 4500, Denton County, Texas; ordinary autozooids, zone of astogenetic change. 2, Holotype, LSU 4500, Denton County, Texas; ordinary autozooids, avicularian polymorphs, and two ovicells, one of which
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Figure8—Differentiation of avicularia in eight species of Wilbertopora plotted against position in cladistic tree. 1, Ratio between distal aperture length and proximal aperture length of avicularia or avicularium-like polymorphs; mean values, W. mutabilis, 0.3246; W. listokinae n. sp., 0.9865; W. tappanae n. sp., 0.9107; W. spatulifera n. sp., 1.1520; W. attenuata n. sp., 1.4621; W. improcera n. sp., 1.1778; W. acuminata n. sp., 1.1230; W. hoadleyae n. sp., 0.8672. 2, Deviation (sum of squared deviations in six avicularian measurements) from mean vector of W. mutabilis

Figure 8 —Differentiation of avicularia in eight species of Wilbertopora ... Available to Purchase

Published: 01 January 2006
Figure 8 —Differentiation of avicularia in eight species of Wilbertopora plotted against position in cladistic tree. 1, Ratio between distal aperture length and proximal aperture length of avicularia or avicularium-like polymorphs; mean values, W. mutabilis , 0.3246; W. listokinae n. sp
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Figure9—Wilbertopora tappanae n. sp., Upper Albian. 1, Paratype, NHM BZ1124 a, Bryan County, Oklahoma, Weno Formation; autozooids in zone of astogenetic change, and autozooids (some with intramural buds) and kenozooid in zone of repetition. 2, Holotype, NHM BZ1124 c, Bryan County, Oklahoma, Weno Formation; ordinary and ovicell-bearing autozooids and kenozooid in zone of repetition. 3, Holotype, NHM BZ1124 c, Bryan County, Oklahoma, Weno Formation; autozooid and avicularium, zone of repetition. 4, Paratype, USNM 526290, HTL-81, Cook County, Texas, Denton Formation; autozooids with spine bases. 5, USNM 526291, Denton County, Texas, Paw Paw Formation; avicularium bearing ovicell on proximal gymnocyst, and autozooids. Scale bar 100 μm

Figure 9 — Wilbertopora tappanae n. sp., Upper Albian. 1, Paratype, NHM... Available to Purchase

Published: 01 January 2006
Figure 9 — Wilbertopora tappanae n. sp., Upper Albian. 1, Paratype, NHM BZ1124 a, Bryan County, Oklahoma, Weno Formation; autozooids in zone of astogenetic change, and autozooids (some with intramural buds) and kenozooid in zone of repetition. 2, Holotype, NHM BZ1124 c, Bryan County, Oklahoma
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Figure10—Wilbertopora spatulifera n. sp., Upper Albian. 1, Paratype, USNM 526305, HTL-96, Williamson County, Texas, Georgetown Formation; zone of astogenetic change and part of zone of repetition, with ordinary autozooids and one avicularium. 2, Holotype, USNM 526292, HTL-9, Grayson County, Texas, Fort Worth Formation; autozooids, ovicells, and avicularia, zone of repetition. 3, Paratype, USNM 526296, HTL-41, Tarrant County, Texas, Fort Worth Formation; “normal” ovicells distal to autozooids and reduced (nonfunctional?) ovicells distal to two avicularia. Scale bars 100 μm

Figure 10 — Wilbertopora spatulifera n. sp., Upper Albian. 1, Paratype,... Available to Purchase

Published: 01 January 2006
Figure 10 — Wilbertopora spatulifera n. sp., Upper Albian. 1, Paratype, USNM 526305, HTL-96, Williamson County, Texas, Georgetown Formation; zone of astogenetic change and part of zone of repetition, with ordinary autozooids and one avicularium. 2, Holotype, USNM 526292, HTL-9, Grayson County
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Figure11—Wilbertopora attenuata n. sp., Upper Albian. 1, Holotype, USNM 526319, HTL-99, Johnson County, Texas, Denton Formation; zone of astogenetic change and part of zone of repetition with ordinary autozooids and two avicularia. 2, Paratype, USNM 526323, HTL-96, Williamson County, Texas, Georgetown Formation; ovicells, maternal autozooids, and two avicularia. Scale bar 100 μm

Figure 11 — Wilbertopora attenuata n. sp., Upper Albian. 1, Holotype, U... Available to Purchase

Published: 01 January 2006
Figure 11 — Wilbertopora attenuata n. sp., Upper Albian. 1, Holotype, USNM 526319, HTL-99, Johnson County, Texas, Denton Formation; zone of astogenetic change and part of zone of repetition with ordinary autozooids and two avicularia. 2, Paratype, USNM 526323, HTL-96, Williamson County, Texas
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Figure12—Wilbertopora improcera n. sp., Upper Albian. 1, Paratype, USNM 526329 HTL-96, Williamson County, Texas, Georgetown Formation; autozooids, zone of astogenetic change. 2, Holotype, USNM 216163, HTL-25, Grayson County, Texas, Denton Formation; ordinary autozooids and two avicularia. 3, Paratype, USNM 216152, HTL-48, Cooke County, Texas, Fort Worth Formation; avicularia, one of which apparently occluded ovicell at its distal margin. 4, Paratype, USNM 526327, HTL-43, Tarrant County, Texas, Fort Worth Formation; maternal and ovicell-bearing autozooids. Scale bar 100 μm

Figure 12 — Wilbertopora improcera n. sp., Upper Albian. 1, Paratype, U... Available to Purchase

Published: 01 January 2006
Figure 12 — Wilbertopora improcera n. sp., Upper Albian. 1, Paratype, USNM 526329 HTL-96, Williamson County, Texas, Georgetown Formation; autozooids, zone of astogenetic change. 2, Holotype, USNM 216163, HTL-25, Grayson County, Texas, Denton Formation; ordinary autozooids and two avicularia
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Figure13—Wilbertopora acuminata n. sp., holotype, USNM 216143, HTL-89, Bell County, Texas, Grayson Formation, Lower Cenomanian. 1, Ordinary autozooids, zone of astogenetic change and autozooids and avicularium, zone of repetition; 2, autozooids and avicularia; ovicells borne on proximal gymnocysts of numerous autozooids and those of two avicularia (upper and lower right). Scale bars 100 μm

Figure 13 — Wilbertopora acuminata n. sp., holotype, USNM 216143, HTL-89,... Available to Purchase

Published: 01 January 2006
Figure 13 — Wilbertopora acuminata n. sp., holotype, USNM 216143, HTL-89, Bell County, Texas, Grayson Formation, Lower Cenomanian. 1, Ordinary autozooids, zone of astogenetic change and autozooids and avicularium, zone of repetition; 2, autozooids and avicularia; ovicells borne on proximal