Relative abundances of conodonts stem from abiotic or biotic causes. High frequencies can result from: 1) biotic positive = high standing crop; 2) biotic negative = lethality (mass mortality); 3) abiotic positive = lag concentrates; 4) abiotic negative = starved sedimentation. Neither abiotic cause should substantially affect the taxonomic composition of the fauna, although either biotic cause—good or bad environmental responses—can and must. Pennsylvanian conodont biofacies are clearly established and evidence of their interrelationships and complexity has continued to mount. We currently recognize no fewer than five levels of conodont biofacies: Ia - Primary generic-level biofacies (examples: Cavusgnathus, Aethotaxis) Ib - Secondary generic-level (“nested”) biofacies (examples: Ellisonia with Cavusgnathus, Hindeodus with Aethotaxis) II - Species-level microbiofacies (examples: Idiognathodus delicatus with Missourian Idioprioniodus / Gondolella , Streptognathodus elegantulus with Missourian Aethotaxis ) III - Apparatus-level biofacies (examples: scottognathoid apparatuses least complete with Cavusgnathus, intermediate with Aethotaxis, most with Idioprioniodus in the Missourian) IV - Ecophenotype variant-level biofacies (examples: perhaps two “species” of Ellisonia with contrasting apparatus plans and morphologies in the Desmoinesian, possible Cavusgnathus morphotypes from the Cavusgnathus- to the Streptognathodus -biofacies).

Abstract: The “Wolfcamp Shale” of the Permian Basin in west Texas is a prime source of oil reserves within the USA. Despite its growing importance, definitive ages and stratigraphic correlations across the basin remain unclear. Fusulinid studies have assigned an approximate age of Late Pennsylvanian (Missourian) to early Permian (Leonardian) for the “Wolfcamp Shale,” but exact horizons for regional stage boundaries are not well defined. Analysis of conodont specimens from three cored basinal shale intervals of the Shell Stevens L1V well in the northern part of the Midland Basin (Lynn County, Texas) improves interpretation of regional stage boundaries within the “Wolfcamp Shale” and provides an accurate means of correlation to depositional cycles on the Eastern Shelf in central Texas. Conodont biostratigraphic markers were compared to log correlations and sequence stratigraphic correlations. Associated gamma-ray (GR) and resistivity (RES) logs show distinct log markers for the Pennsylvanian–Permian boundary and base of the Leonardian Stage that are traceable to immediately adjacent wells and possibly to portions of the southern part of the Midland Basin. Third- and fourth-order sequence stratigraphic cycles were delineated using core lithology, conodont abundances, and gamma-ray and resistivity readings. These sequences are linked to those observed in the Eastern Shelf stratigraphic sections and provide a useful secondary mode of correlation.

Abstract Late Pennsylvanian conodont faunas were dominated by idiognathodids historically assigned to Idiognathodus (flat P 1 ) or Streptognathodus (troughed P 1 ). Recent work suggests clades arose iteratively, through time, from unrelated ancestors in different geographical regions. The end-Desmoinesian extinction event terminated two major genera, Swadelina (troughed) and Neognathodus (long carina), and comparable new morphotypes developed from surviving Idiognathodus species in the early Kasimovian, especially in North America. True Streptognathodus (troughed) and Heckelina n. gen (asymmetric, eccentric groove) appeared in North America in the mid-Kasimovian. Another troughed clade arose in Eurasia (‘ S. ’ 2) and attained a global distribution by the late Kasimovian. A second, early Gzhelian, Eurasian radiation produced new troughed forms (‘ S. ’ 4) that dominated Gzhelian faunas globally. In South China, endemic clades of eccentrically grooved Idiognathodus ? and troughed forms (‘ S .’ 3) appeared in the late Kasimovian and persisted into the Gzhelian. Typical Idiognathodus species were uncommon by the late Kasimovian and disappeared in the mid-Gzhelian. After a low diversity interval in the mid-Gzhelian, a new major radiation of weakly troughed forms occurred (‘ S. ’ 5), which led to redevelopment of Idiognathodus -like elements in the Cisuralian. Other conodont genera from offshore ( Gondolella, Idioprioniodus ) and nearshore settings ( Hindeodus, Diplognathodus, Adetognathus, Ellisonia ) are poorly studied and show low diversity and little morphological change.

Abstract Carboniferous conodont biostratigraphy comprises regional zonations that reflect the palaeogeographical distribution of taxa and distinct shallow-water and deep-water conodont biofacies. Some species have a global distribution and can effect high quality correlations. These taxa are incorporated into definitions of global Carboniferous chronostratigraphic units. A standard global Carboniferous zonation has not been developed. The lowermost Mississippian is zoned by Siphonodella species, excepet in shallow-water facies, where other polygnathids are used. Gnathodus species radiated during the Tournaisian and are used to define many Mississippian zones. A late Tournaisian maximum in diversity, characterized by short-lived genera, was followed by lower diversity faunas of Gnathodus species and carminate genera through the Visean and Serpukhovian. By the late Visean and Serpukhovian, Lochriea provides better biostratigraphic resolution. Shallow-water zonations based on Cavusgnathus and Mestognathus are difficult to correlate. An extinction event near the base of the Pennsylvanian was followed by the appearance of new gnathodid genera: Rhachistognathus , Declinognathodus , Neognathodus , Idiognathoides and Idiognathodus . By the middle of the Moscovian, few genera remained: Idiognathodus , Neognathodus and Swadelina. During the middle Kasimovian and Gzhelian, only Idiognathodus and Streptognathodus species were common. Near the end of the Gzhelian, a rediversification of Streptognathodus species extended into the Cisuralian.

Abstract An insect zonation with a time resolution of 1.5–2 Ma for Late Pennsylvanian to Early Permian (Kasimovian to Artinskian) non-marine deposits is presented. The zonation is based on the directed morphogenetic evolution of colour pattern in the forewings of the blattid (cockroach) family Spiloblattinidae. This evolution is observed in lineages of succeeding species of three genera. All three genera are widely distributed in the palaeo-equatorial zone from Europe to North America, that is, in the Euramerican biota province. Increasing reports of spiloblattinid zone species in condont-bearing, interfingered marine/continental strata of North American Appalachian, Mid-Continent and West Texas basins could be the key to direct biostratigraphical correlations of pure continental profiles, as are present in the most parts of the Hercynides, to the global marine scale.

A thick succession of upper Paleozoic carbonate rocks and minor chert crops out north of the head of Otto Fiord (northwest [NW] Ellesmere Island, Nunavut) in the Canadian Arctic Archipelago. These rocks accumulated in a tectonic subbasin—the Otto Fiord Depression (OFD)—of the Sverdrup Basin that likely originated through rifting during late Early Carboniferous (Serpukhovian). Following a long interval of passive subsidence that allowed a thick succession of Moscovian–Kasimovian carbonate rocks to fill the OFD, tectonic activity resumed during the Gzhelian (Late Pennsylvanian). This resulted in rapid collapse of the depression along its axis and simultaneous uplifts of its margins, a style of tectonism in accord with the inferred basin-wide shift to a transpressional–transtensional stress regime at that time. Late Pennsylvanian–Early Permian sedimentation in the OFD led to the development of four long-term (second-order) transgressive–regressive sequences of early Gzhelian–middle Asselian (<1200 m), late Asselian–late Sakmarian (<380 m), latest Sakmarian–late Artinskian (<160 m) and latest Artinskian–late Kungurian (<60 m) age. These ages are supported by integration of biostratigraphic data from conodonts, fusulinaceans, and small foraminifers. The development of each sequence-bounding unconformity was associated with renewed tectonism in the OFD. Each sequence recorded the development of a depositional system characterized by high energy peripheral shoreface grainstones passing basinward across a gently dipping ramp into deep-water basinal calcareous and siliceous mudstone. The ramp portion of the early Gzhelian–middle Asselian system comprises both cool-heterozoan to warmphotozoan carbonates (Nansen Formation) suggesting a relatively shallow thermocline at that time. These rocks are arranged in a series of high-order cyclothems of glacio-eustatic origin. Cyclothemic sedimentation ended at the Asselian–Sakmarian boundary, simultaneous to a major depositional system shift to cool-water heterozoan sedimentation (Raanes Formation), a change presumably brought on by the closure of the Uralian seaway linking NW Pangea with the Tethyan Ocean. This event led to the destruction of the permanent thermocline, and disappearance of photozoan carbonates by the early Sakmarian despite rising temperatures globally. Cool-water heterozoan sedimentation, associated with relatively shallow outer-ramp to midramp spiculitic chert resumed in the Artinskian and then again in the Kungurian (Great Bear Cape Formation) when the OFD was filled up. The depression ceased to exist as a separate tectonic/subsidence entity with the widespread sub-Middle Permian unconformity, above which sediments were deposited during a passive subsidence regime across most of the Sverdrup Basin. The Pennsylvanian–Lower Permian succession that accumulated in the OFD along the clastic-free northern margin of the Sverdrup Basin is essentially identical, both in terms of tectonic evolution and stratigraphic development, with the coeval succession of Raanes Peninsula, southwest (SW) Ellesmere Island, the type area of the Raanes, Trappers Cove, and Great Bear Cape formations along the clastic-influenced southern margin.

Abstract We present an updated look at Carboniferous brachiopod biozonation from most of the world framed into a revised Carboniferous palaeogeography, based on a selection of the literature published on Carboniferous brachiopods since the nineteenth century. The biostratigraphic significance of the most important brachiopod taxa is synthesized in seven geographical correlations. The Mississippian is characterized by rich brachiopod faunas, with widespread taxa with a good potential for global correlation, such as Rugosochonetes , Delepinea , Buxtonia , Antiquatonia , Spinocarinifera , Marginatia , Fluctuaria , Ovatia , Rhipidomella , Lamellosathyris , Unispirifer , Tylothyris and Syringothyris . From the mid-Visean to the late Serpukhovian, taxa of gigantoproductidines are biostratigraphically significant, and occur everywhere except South America and Australia, which remain as distinct faunal successions for most of the period. A major turnover occurs at the beginning of the Pennsylvanian, characterized by a higher degree of provincialism. Pennsylvanian brachiopod faunas are diverse in China, Russia and North America, but otherwise they are less developed and are characterized mostly by endemic taxa, hampering long-distance correlation. An exception is the rapid diversification of taxa of the Choristitinae, which were widespread from the Bashkirian to the Moscovian, allowing long-distance correlation.