Abstract: A section approximately 30 m thick in the uppermost part of the deltaic lower Pleistocene Tursi Sandstone at Rabatana has been analyzed. It contains Bichordites monastiriensis, rare Macaronichnus segregatis, and very rare Rosselia socialis (Bichordites ichnoassemblage) in the lower part, and abundant B. monastiriensis and R. socialis, and rare M. segregatis ( Bichordites-Rosselia ichnoassemblage), in the upper part. Ichnofabric and sedimentologic features indicate upper- to middle-shoreface conditions in the lower part of the section, decreasing in energy up section to lower-shoreface conditions. The Bichordites ichnoassemblage is related to unstable conditions on the sea floor, which favored infaunal colonization by vagile irregular echinoids. The Bichordites-Rosselia ichnoassemblage is related to more stable conditions in a probably deeper and / or protected environment, and to clearer waters in a declining deltaic system. Under such conditions, colonization by sedentary ?terebelloid tracemakers of R. socialis, which fed on the sea floor, was possible. The low trace-fossil diversity is related to intense bioturbation by echinoids, which prevented colonization by other tracemakers, except for the Rosselia tracemakers.

Abstract Recent work has focused on erecting new Seilacherian ichnofacies for depositional environments subject to recurring temporal and spatial variations in physico-chemical stress. In marine deltaic settings, these correspond to the Phycosiphon Ichnofacies for mudstone-dominated prodeltaic deposits and the Rosselia Ichnofacies for sandstone-dominated delta-front successions. The archetypal expressions of these ichnofacies, however, are founded on mixed process (wave- and river-influenced) systems, because the juxtaposition of ambient marine conditions during periods of prolonged wave energy with rapid deposition and physico-chemically stressed conditions during heightened fluvial discharge best expresses the deltaic signal. As deltaic settings shift towards end-member processes (e.g. river domination, wave domination and tide domination), or towards mixed-process conditions other than river and wave influence, the resulting ichnological suites and bioturbation fabrics depart from the recently published archetypes. Using selected studies of marine deltaic deposits, predictable departures from the archetypes can be recognized on the basis of these changing processes and their associated physico-chemical stresses. River-dominated delta deposits and tide-dominated delta successions display the greatest deviation from the published archetypes. River-dominated examples show elevated deposition rates, periods of salinity reduction, slumping and dewatering, elevated water turbidity, flood-induced sediment gravity flows and hypopycnal-generated fluid mud. As a result, river-dominated successions are largely devoid of bioturbation. Evidence of marine conditions is commonly restricted to isolated occurrences of dwelling structures such as Arenicolites , Ophiomorpha or Rosselia in sandstone, and Chondrites , Phycosiphon or Zoophycos in mudstone beds, particularly in prodeltaic intervals. Tide-dominated deltaic successions are markedly heterolithic and typified by highly mobile substrates manifested by incrementally migrating asymmetric bedforms and abundant fluid mud. Such settings are also prone to marked changes in salinity and shifts in the position of the turbidity maximum zone. Successions typically show low intensities of bioturbation and sporadically distributed burrows, as well as deposit-feeding structures, deeply penetrating dwelling structures or fugichnia. Many trace fossil suites consist entirely of facies-crossing elements, making assignment to an ichnofacies impossible. Storm flood-dominated deltaic successions are characterized by tempestites that are typically interstratified with river flood-induced sediment-gravity flow deposits and/or mantled by largely unburrowed mudstone drapes derived from hypopycnal plumes associated with river floods. Where these storm flood cycles are interstratified with ambient fairweather beds, assignment to the archetypal deltaic ichnofacies is straightforward. However, as storm beds become increasingly erosionally amalgamated, the preservation potential of the fairweather beds is reduced and the resulting trace fossil suites are biased towards those recording opportunistic colonization of the event beds. The presence of mudstone layers with low bioturbation intensity (BI) containing small numbers of ichnogenera positively correlated with marine conditions (e.g. Chondrites , Phycosiphon and/or Zoophycos ) may be the only evidence that the suites should be assigned to one of the deltaic ichnofacies. Wave-dominated deltas lacking significant storm influence are typically challenging to differentiate from their archetypal strandplain shoreface counterparts and, correspondingly, the resulting trace fossil suites are broadly comparable to the archetypal Cruziana and Skolithos ichnofacies. Most of the preserved record of wave-dominated delta successions is related to fairweather ambient conditions, and so facies typically show high BI values and uniformly distributed bioturbation. Key to recognizing that the suites should be assigned to one of the deltaic ichnofacies is the presence of rare river-generated mudstone and sandstone beds that display evidence of physico-chemical stress and/or the paucity of domichnia typical of suspension-feeding organisms. In most delta types, the prodeltaic facies are most readily discerned to contain trace fossil suites of the Phycosiphon Ichnofacies, owing to the higher preservation potential of all depositional processes, including marine fairweather beds, river-supplied hyperpycnites and other sediment gravity flow deposits, tempestites and fluid mud derived from river flood-related hypopycnal plumes. Assignment of trace fossil suites to the Rosselia Ichnofacies requires some record of the fairweather conditions, which are generally diminished in river-, tide- and storm-dominated successions. The dominance of structures positively correlated to deposit-feeding ethologies at the expense of those attributed to suspension-feeding strategies may point to elevated water turbidity and assignment of the suite to the Rosselia Ichnofacies. However, in many cases, the ichnological suites of delta fronts are so depauperate that assignment to an ichnofacies is problematic and should be avoided.