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Pterodactyloidea
New material of dsungaripterid pterosaurs (Pterosauria: Pterodactyloidea) from western Mongolia and its palaeoecological implications
Early juvenile pterosaurs (Pterodactyloidea: Pterodaustro guinazui ) from the Lower Cretaceous of central Argentina
Pterosaur (Pteranodontoidea, Pterodactyloidea) scapulocoracoid from the Early Cretaceous of Venezuela
Abstract The discovery of a left scapula and coracoid (MBLUZ P-911) representing the first evidence of a pterosaur from Venezuela is reported here. The material comes from the Lower Cretaceous (Aptian) Apón Formation, in the northwestern part of the country. In MBLUZ P-911 the scapula is significantly smaller than the coracoid, a synapomorphy of the Pteranodontoidea, according to Kellner. The coracoid of the Venezuelan specimen is more elongated and gracile than those of Istiodactylus and Pteranodon, and also lacks the ventromedial coracoidal flange present in the latter. Overall MBLUZ P-911 is very similar to the scapulocoracoid of the Anhangueridae, including the presence of a longitudinal ridge on the medial surface of the coracoid and a comparatively short scapula, and is therefore tentatively referred to this taxon. This occurrence extends the pterosaur record to the northern part of the South American portion of Gondwana.
Abstract This paper deals with the systematics of tetrapod ichnotaxa based on footprints. Beyond the consideration of the nomenclatural rules for ichnotaxa in the ICZN, this paper tries to determine how to establish an ichnotaxonomy that reflects the identity of the track-maker (the organism that has made the track) and how to include this ichnotaxonomy in the skeleton-based taxonomy. This leads to the establishment of several criteria, e.g. ichnospecies should be defined on the print morphology and the relative position of the prints (including the variability due to the track-maker's dynamics), two ichnospecies should represent different species, the ichnospecies and ichnogenus levels are sufficient to discriminate the ichnotaxa and link them to the skeleton-based taxonomy. These ichnotaxonomical criteria are applied to a revision of the ichnogenus Pteraichnus Stokes 1957 (Pterosauria, Pterodactyloidea). Only the type species Pteraichnus saltwashensis is considered as valid, the pterosaurian origin of Purbeckopus is questioned and Agadirichnus is highlighted because it could be a senior synonym of Pteraichnus. The result of this drastic revision underlines the importance of the proposed ichnotaxonomical principles to avoid the unfounded proliferation of tetrapod ichnotaxa.
On the taxonomic status of Cycnorhamphus and Gallodactylus (Pterosauria; Pterodactyloidea)
Pterosaur phylogeny and comments on the evolutionary history of the group
Abstract A cladistic analysis based on 39 terminal taxa and 74 characters (several multistate) using PAUP (Phylogenetic Analysis Using Parsimony) (3.1.1 for Macintosh and 4.0b10 for Microsoft Windows) presents a new hypothesis of pterosaur inter-relationships. This study suggests that the most primitive taxon is the Anurognathidae, followed by Sordes and all remaining pterosaurs. Dendrorhynchoides is confirmed as a member of the Anurognathidae, being closely related to Batrachognathus. Preondactylus occupies a more derived position than Sordes, which questions its previous assignment as the most primitive pterosaur. The hypothesis of rhamphorhynchoid paraphyly is confirmed, with the Rhamphorhynchidae more closely related to the Pterodactyloidea than to more basal forms. The Pterodactyloidea shows a basal dichotomy: the Archaeopterodactyloidea and the Dsungaripteroidea. The Archaeopterodactyloidea is formed by Pterodactylus + Germanodactylus and a clade formed by Gallodactylidae + Ctenochasmatidae. The Nyctosauridae occupies the basal position within dsungaripteroids and is followed by the Pteranodontoidea and the Tapejaroidea. Pteranodontoids have Pteranodon at the base, followed stepwise by Istiodactylus, Ornithocheirus and the Anhangueridae. Tapejaroids are composed of the Dsungaripteridae at the base followed by the Tapejaridae and the Azhdarchidae. Major trends within pterosaur evolutionary history are: general increase in size (wing span and body); increase of wing metacarpal and pteroid; decrease of proportional length of the second and third wing phalanx relative to the first; gradual increase of rostrum (anterior to external nares); and anterior shift of the skull-mandible articulation. Cranial crests are present in most pterodacty-loids, but markedly in the Ornithocheiroidea, where all taxa show some sort of crest on the skull. The loss of teeth, previously assumed to have occurred independently in several lineages, seems to be a general trend among dsungaripteroids. Several nodes recovered by this analysis are supported by very few characters, a result at least partially attributable to the limited available information from several taxa due to poor preservation and/or preparation.
Abstract Previous cladistic studies of pterosaur relationships suffer from restricted numbers of taxa and characters, incomplete data sets and absence of information on characters, tree structure and the robustness of trees. Parsimony analysis of a new character data set (60 characters, 20 terminal taxa, 93.75% complete) yielded six trees. In the strict consensus tree Preondactylus is the most basal taxon followed, stepwise, by the Dimorphodontidae and the Anurognathidae. Beyond this basal group, more derived pterosaurs (Campylognathoididae (Rhamphorhynchidae + Pterodactyloidea)) share a suite of characters principally associated with elongation of the rostrum. The Pterodactyloidea consists of four major clades. The Ornithocheiroidea is the most basal taxon consisting, stepwise, of Istiodactylus, the Ornithocheiridae, Nyctosaurus and the Pteranodontidae. The remaining taxa, Ctenochasmatoidea, Dsungaripteroidea and Azhdarchoidea, are weakly united in a clade of non-ornithocheiroid pterodactyloids, but their inter-relationships are difficult to resolve. Cycnorhamphus is the basal-most ctenochasmatoid, while the remaining taxa (Pterodactylus, Lonchodectidae, Ctenochasmatidae) form an unresolved trichotomy. The Dsungaripteroidea (Germanodactylus + Dsungaripteridae) is strongly supported by features of the skull and dentition. The Azhdarchoidea (Tapejara [Tupuxuara + Azhdarchidae]) is united by cranial characters such as elevation of the antorbital region, and relative shortening of the wing finger. The pattern of pterosaur evolution suggested by the results of this analysis is broadly similar to traditional ideas, but has greater resolution, more complexity and reveals several previously unrecognized ‘events’.
Abstract Two specimens of a tapejarid pterosaur (Pterodactyloidea, Tapejaridae) are described as representing a new species. Both specimens show evidence for soft tissues preserved in association with a sagittal bony cranial crest. Both specimens are from the Nova Olinda Member Konservat Lagerstätte of the Crato Formation of the Araripe Basin, northeastern Brazil. They represent the second tapejarid species from this formation. Comparisons are made with other crested pterosaurs and comments on the utility and aerodynamics of pterosaurian head crests are made.