The Bird Spring Shelf in southeastern California, along with coeval turbidite basins to the west, records a complex history of late Paleozoic sedimentation, sea-level changes, and deformation along the western North American continental margin. We herein establish detailed correlations between deposits of the shelf and the flanking basins, which we then use to reconstruct the depositional history, paleogeography, and deformational history, including Early Permian emplacement of the regionally significant Last Chance allochthon. These correlations are based on fusulinid faunas, which are numerous both on the shelf and in the adjoining basins. Study of 69 fusulinid species representing all major fusulinid-bearing Pennsylvanian and Lower Permian limestone outcrops of the Bird Spring Shelf in southeastern California, including ten new species of the genera Triticites , Leptotriticites , Stewartina , Pseudochusenella , and Cuniculinella , forms the basis for our correlations. We group these species into six fusulinid zones that we correlate with fusulinid-bearing strata in east-central and southern Nevada, Kansas, and West Texas, and we propose some regional correlations not previously suggested. In addition, we utilize recent conodont data from these areas to correlate our Early Permian fusulinid zones with the standard Global Permian Stages, strengthening their chronostratigraphic value. Our detailed correlations between the fusulinid-bearing rocks of the Bird Spring Shelf and deep-water deposits to the northwest reveal relationships between the history of shelf sedimentation and evolution of basins closer to the continental margin. In Virgilian to early Asselian (early Wolfcampian) time (Fusulinid Zones 1 and 2), the Bird Spring Shelf was flanked on the west by the deep-water Keeler Basin in which calcareous turbidites derived from the shelf were deposited. In early Sakmarian (early middle Wolfcampian) time (Fusulinid Zone 3), the Keeler Basin deposits were uplifted and transported eastward on the Last Chance thrust. By middle Sakmarian (middle middle Wolfcampian) time (within Fusulinid Zone 4), emplacement of the Last Chance allochthon was complete, and subsidence caused by thrust loading had resulted in development of a new turbidite basin (Darwin Basin) along the former western part of the Bird Spring Shelf. At the same time, farther east into the craton, paralic facies began prograding westward, so that the youngest fusulinid-bearing limestones on the shelf in this area become progressively younger to the west. Eventually, in Artinskian to Kungurian (late Wolfcampian to Leonardian) time (Fusulinid Zones 5 and 6), deposition of fusulinid-bearing limestone on the shelf was restricted to a marginal belt between the prograding paralic facies to the east and the Darwin Basin to the west. Development of the Keeler Basin in Pennsylvanian to earliest Permian time was approximately coeval with collision between South America-Africa (Gondwana) and North America (Laurentia) on the Ouachita-Marathon orogenic belt. This basin developed inboard of a northwest-trending, sinistral fault zone that truncated the continental margin. Later, in the Early Permian, the Last Chance allochthon, which was part of a northeast-trending belt of deformation that extended into northeastern Nevada, was emplaced. This orogenic belt probably was driven by convergence at the continental margin to the northwest. This work adds significant detail to existing interpretations of the late Paleozoic as a time of major tectonic instability on the continental margin of southeastern California as it changed from a relatively passive margin that had characterized most of the Paleozoic to an active convergent margin that would characterize the Mesozoic.

Abstract: A review of Permian fusuline biostratigraphy is made in this paper in order to improve the correlation of Permian strata globally. Permian fusuline biostratigraphy in the Tethyan and Panthalassan regions can be correlated roughly because the fusulines had good faunal communications between these two regions. However, fusuline faunas from the North American Craton region were devoid of almost all neoschwagerinids and dominated exclusively by schwagerinids during the Guadalupian (Middle Permian) because of the blockage caused by the vast Pangaea supercontinent. This renders the correlation of Middle Permian biostratigraphy and chronostratigraphy between the Tethyan region and North American region challenging. Significant evolutionary key points in fusulines include the first occurrence of Pseudoschwagerina or Sphaeroschwagerina during the earliest Permian, first occurrence of Pamirina and Misellina during the Yakhtashian and Bolorian, and the extinction of all schwagerinids and neoschwagerinids by the end of the Midian.

Abstract This paper proposes a synthesis of the taxonomy, phylogeny, palaeogeographic distribution, regional biostratigraphy, and palaeobiogeographic faunal development of Carboniferous fusuline foraminifers. They appeared in the latest Tournaisian and comprised a small-sized, morphologically conservative taxonomic group during the Mississippian. Fusulines became larger and prevailed in Pennsylvanian foraminiferal assemblages. Carboniferous fusulines consist of Ozawainellidae, Staffellidae, Schubertellidae, Fusulinidae, and Schwagerinidae, in which 95 genera are considered as valid taxonomically. Upsizing their shells throughout the Pennsylvanian is likely related to symbiosis with photosynthetic microorganisms, which was accelerated by the acquisition of a keriothecal wall in Late Pennsylvanian schwagerinids. Regional fusuline succession data from 40 provinces provide a refined biostratigraphy, enabling zonation and correlation with substage- or higher-resolution precision in the Pennsylvanian. Their spatio-temporal faunal characteristics show that fusulines had a cosmopolitan palaeobiogeographic signature in Mississippian time, suggesting unrestricted faunal exchange through the palaeoequatorial Rheic Ocean. After the formation of Pangaea, Pennsylvanian fusulines started to show provincialism, and their distributions defined the Ural–Arctic Region in the Boreal Realm, Palaeotethys, Panthalassa, and North American Craton regions in the Palaeoequatorial Realm, and Western Gondwana and Eastern Peri-Gondwana regions in the Gondwana Realm. The Western Palaeotethys and East European Platform Subregions maintained higher generic diversity throughout the Pennsylvanian.