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Journal Article

THE UPPER GIVETIAN (MIDDLE DEVONIAN) SUBTERMINUS CONODONT ZONE IN NORTH AMERICA, EUROPE AND NORTH AFRICA

KATARZYNA NARKIEWICZ, PIERRE BULTYNCK
Published: 01 July 2010
Journal of Paleontology (2010) 84 (4): 588–625.
DOI: https://doi.org/10.1666/10-029.1
.... The holotype of Icriodus subterminus from the North Liberty beds in Iowa is most likely a specimen that was reworked from the Cedar Valley Limestone. Icriodus cedarensis and Icriodus tafilaltensis are described as new species, and the diagnoses of Icriodus excavatus and Icriodus expansus are amended. Between...
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Figure7—All elements from Rapid Mb., Conklin Quarry, Iowa (USA): all figures are P1 elements; scale bars are 100 µm. 1, 2, 3, Icriodus subterminus Youngquist, α morphotype, upper, lateral and lower views, SUI 100607 117328; 4–13, Icriodus subterminus Youngquist, β morphotype; 4, 5, 6, upper, lateral and lower views, SUI 100609 117337; 7, 8, 9, upper, lateral and lower views, SUI 100608 117332; 10, 11, upper and lateral views, SUI 100607 117329; 12, 13, upper and lateral views, SUI 100608 117333; 14–15, 21–26, Icriodus cedarensis n. sp.; 14, 15, upper and lateral views of paratype, SUI 100608 117334; 21, 22, 23, upper, lateral and lower views of holotype, SUI 100609 117338; 24, 25, 26, upper lateral and lower views, SUI 100607 117330; 16–17, Icriodus aff. I. subterminus Youngquist, upper and lateral views, SUI 100608 117335; 18, 19, 20, Icriodus excavatus Weddige, upper, lateral and lower views, SUI 100608 117336; 27, Icriodus tafilaltensis n. sp. upper view, SUI 100607 117331.

Figure 7 —All elements from Rapid Mb., Conklin Quarry, Iowa (USA): all fig...

Published: 01 July 2010
; 18 , 19 , 20 , Icriodus excavatus Weddige, upper, lateral and lower views, SUI 100608 117336; 27 , Icriodus tafilaltensis n. sp. upper view, SUI 100607 117331.
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Figure18—at the Mehaffey Bridge section and Figs. 7–19 from Rapid Mb. in the Solon Quarry, Iowa, (USA): all figures are P1 elements; scale bars are 100 µm. 1, 2, Icriodus subterminus Youngquist, β morphotype, upper and lateral views, sample Iowa PB-1972-2, IRScNB b5254. 3, 4, Pandorinellina insita (Stauffer), lateral and upper views, sample Iowa PB-1972-2, IRScNB b5255. 5, 6, Icriodus subterminus Youngquist, α morphotype, upper and lateral views, sample Iowa PB-1972-2, IRScNB b5256. 7–9, 13–16, Icriodus expansus Branson and Mehl: 7, 8, 9, upper, lateral and lower views, sample Iowa PB-1972-3, IRScNB b5257; 13, upper view, sample Iowa PB-1972-3, IRScNB b5258; 14, 15, 16, upper, lateral and lower views; sample Iowa PB-1972-3, IRScNB b5259. 10, 11, 12, Icriodus tafilaltensis n. sp.: 10, upper view, sample Iowa PB-1972-3, IRScNB b5260; 11, 12, upper and lateral views of paratype, sample Iowa PB-1972-3, IRScNB b5261. 17, Icriodus brevis Stauffer, upper view, sample Iowa PB-1972-3, IRScNB b5262. 18, 19, Icriodus difficilis Ziegler and Klapper, upper and lateral views, sample Iowa PB-1972-3, IRScNB b5263.

Figure 18 —at the Mehaffey Bridge section and Figs. 7–19 from Rapid Mb. ...

Published: 01 July 2010
views, sample Iowa PB-1972-3, IRScNB b5257; 13 , upper view, sample Iowa PB-1972-3, IRScNB b5258; 14 , 15 , 16 , upper, lateral and lower views; sample Iowa PB-1972-3, IRScNB b5259. 10 , 11 , 12 , Icriodus tafilaltensis n. sp.: 10 , upper view, sample Iowa PB-1972-3, IRScNB b5260; 11 , 12
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Figure15—All elements are from Cambresèque Mb. of the Beaulieu Fm.; BN samples are from the Banc Noir Quarry, others are from the Caffiers/Ferques section, Boulonnais (France); except Fig. 6 all figures are P1 elements; scale bars are 100 µm. 1, 2, Icriodus tafilaltensis n. sp., upper and lateral views, Cédule Limestone, sample BN1, IRScNB b5214. 3–6, Ancyrodella pristina Khalymbadzha and Chernysheva, Cédule Lmst., sample BN1: 3, lower view, IRScNB b5215; 4, 5, upper and lower views, IRScNB b5216; 6, lateral view of P2 element, IRScNB b5217. 7, 8, Ancyrodella binodosa Uyeno, upper and lower views, Cédule Lmst., sample BN3 IRScNB b5218. 9, 10, 13–16, Icriodus cedarensis n. sp.: 9, 10, upper and lateral views, Cédule Lmst., sample BN1, IRScNB b5219; 13, 14, upper and lateral views, Cédule Lmst., sample BN3, IRScNB b5220; 15, 16, upper and lateral views, Cambresèque Mb., sample 32, IRScNB B5221. 11, 12, Icriodus symmetricus Branson and Mehl, upper and lateral views, Cédule Lmst., sample BN1, IRScNB b5222. 17, 18, Icriodus excavatus Weddige, upper and lateral views, Cambresèque Mb., sample 32, IRScNB b5223. 19, 20, Icriodus subterminus Youngquist α morphotype, upper (reillustration from Brice et al., 1979a, pl. 27, fig. 26, IRScNB b1132) and lateral views, Cambresèque Mb., sample 32, IRScNB b1132. 21, 22, Icriodus subterminus Youngquist, β morphotype, upper and lateral views, Cambresèque Mb., sample 32, IRScNB b5224. 23, 24, Icriodus aff. I. cedarensis, upper and lateral views, Cambresèque Mb., sample 32, IRScNB b5225.

Figure 15 —All elements are from Cambresèque Mb. of the Beaulieu Fm.; BN ...

Published: 01 July 2010
Figure 15 —All elements are from Cambresèque Mb. of the Beaulieu Fm.; BN samples are from the Banc Noir Quarry, others are from the Caffiers/Ferques section, Boulonnais (France); except Fig. 6 all figures are P 1 elements; scale bars are 100 µm. 1 , 2 , Icriodus tafilaltensis n. sp., upper
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Figure19—All elements are from Bou Dîb Fm. of the Ma'der area (E Anti-Atlas, Morocco); all figures are P1 elements; scale bars are 100 µm. 1, 2, Schmidtognathus wittekindti Ziegler, upper and lateral views, Bou Dîb section, Upper Member, sample 6, IRScNB b5271. 3, Schmidtognathus peracutus (Bryant), upper view, Bou Dîb section, Upper Mb., sample 5, IRScNB b5272. 4, 5, Icriodus cedarensis n. sp., upper and lateral views, Bou Dîb section, Upper Mb., sample 6, IRScNB b5273. 6, Klapperina disparalvea (Orr and Klapper), upper view, Hillock 760-Bou Terga section, Upper Mb., sample 12, IRScNB b5274. 7, 8, Polygnathus cristatus ectypus Huddle, upper and lower views, Hillock 760–Bou Terga section, Middle Mb., sample 7, IRScNB b5275. 9, 10, Polygnathus ovatinodosus, Ziegler and Klapper, upper and lateral views, Hillock 760–Bou Terga section, Middle Mb., sample 7, IRScNB b5276. 11, 12, Icriodus tafilaltensis n. sp., upper and lower views, Hillock 760–Bou Terga section, Upper Mb., sample 12, IRScNB b5277. 13, Icriodus difficilis Ziegler and Klapper, upper view, Hillock 760–Bou Terga section, Upper Mb., sample 12, IRScNB b5278. 14, 15, Polygnathus pollocki Druce, upper and lower views, Hillock 760-Bou Terga section, Upper Mb., sample 15, IRScNB b5279.

Figure 19 —All elements are from Bou Dîb Fm. of the Ma'der area (E Anti-A...

Published: 01 July 2010
, IRScNB b5276. 11 , 12 , Icriodus tafilaltensis n. sp., upper and lower views, Hillock 760–Bou Terga section, Upper Mb., sample 12, IRScNB b5277. 13 , Icriodus difficilis Ziegler and Klapper, upper view, Hillock 760–Bou Terga section, Upper Mb., sample 12, IRScNB b5278. 14 , 15 , Polygnathus
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Figure17—All elements are from Bouia Fm. of Achguig Group, from Bou Tchrafine section BT I of the Tafilalt area (E Anti-Atlas, Morocco); all figures are of P1 elements; scale bars are 100 µm. 1–7, 12, 13, Icriodus tafilaltensis n. sp., sample 33b: 1, 2, 3, upper, lateral and lower views of holotype, IRScNB b5241; 4, 5 and 6, 7, upper and lateral views of paratypes IRScNB b5242 and b5243, 12, 13, upper and lower views of paratype IRScNB b5244. 8, Ozarkodina semialternans (Wirth), lateral view, sample 31b, IRScNB b5245. 9, 10, Icriodus expansus Branson and Mehl, upper and lateral views, sample 33b, IRScNB b5246. 11, Icriodus difficilis Ziegler and Klapper, upper view, sample 43, IRScNB b5247. 14, 15, Polygnathus cristatus ectypus Huddle, upper and lower views, sample 35, IRScNB b5248. 16, 17, Polygnathus dengleri dengleri Bischoff and Ziegler, lateral and upper views, sample 40, IRScNB b5249. 18, Schmidtognathus wittekindti Ziegler, upper view, sample 41, IRScNB b5250. 19, Schmidtognathus latifossatus Wirth, upper view, sample 33b, IRScNB b5251. 20, 21, Icriodus subterminus Youngquist, β morphotype, upper and lateral views (reillustration from Bultynck and Hollard, 1980, pl. 8, fig. 22, IRScNB b1011), sample 37. 22, 23, Icriodus cedarensis n. sp., upper and lateral views (reillustration from Bultynck, 2003, pl. 3, fig. 22, IRScNB b4237), sample 37. 24, 25, Ancyrodella cf. A. pristina Khalymbadzha and Chernysheva, upper and lower views, sample 37, IRScNB b5252. 26, 27, Pandorinellina insita (Stauffer), upper and lateral views, sample 43, IRScNB b5253.

Figure 17 —All elements are from Bouia Fm. of Achguig Group, from Bou Tchr...

Published: 01 July 2010
Figure 17 —All elements are from Bouia Fm. of Achguig Group, from Bou Tchrafine section BT I of the Tafilalt area (E Anti-Atlas, Morocco); all figures are of P 1 elements; scale bars are 100 µm. 1–7 , 12 , 13 , Icriodus tafilaltensis n. sp., sample 33b: 1 , 2 , 3 , upper, lateral
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Figure16—All elements are from Bouia Fm. of Achguig Group, from Bou Tchrafine sections BT II and BT III of the Tafilalt area (E Anti-Atlas, Morocco); except Figs. 20, 21 all figures are P1 elements; scale bars are 100 µm. 1, 2, Polygnathus dengleri dengleri Bischoff and Ziegler, upper and lateral/oblique views, BT II section, sample 10, IRScNB b5228. 3, 22, Polygnathus webbi Stauffer: 3, upper view, BT II section, sample 12, IRScNB b5226; 22, upper view, BT III, sample 1, IRScNB b5227. 4, 5, Icriodus subterminus Youngquist, α morphotype, upper and lateral views, BT II section, sample 12, IRScNB b5229. 6, 7, Icriodus subterminus Younquist, β morphotype, upper (reillustration from Bultynck, 1986, pl. 2, fig. 14, IRScNB b1800) and lateral views, BT II section, sample 12, IRScNB b1800. 8, 19, 20, 21, Skeletognathus norrisi (Uyeno): 8, upper view, BT II section, sample 12, IRScNB b5231; 19, upper view, BT III, sample 1, IRScNB b5232; 20, 21, upper and lateral views of P2 element, BT III, sample 1, IRScNB b5233. 9, Mesotaxis falsiovalis/ovalis→Palmatolepis transitans Müller, upper view, BT II sample 14, IRScNB b5234. 10, 11, 12, Icriodus tafilaltensis n. sp., upper, lateral and lower views, BT II, sample 12, IRScNB b5235. 13, 14, Polygnathus pollocki Druce, upper and lateral views, BT II, sample 10, IRScNB b5236. 15, Palmatolepis transitans Müller, upper view, BT II, sample 14, IRScNB b5237. 16, 17, 18, Mesotaxis falsiovalis Sandberg, Ziegler and Bultynck/M. ovalis (Ziegler and Klapper), upper and lower views and enlargement of the pit, BT III, sample 8, IRScNB b5238. 23, Pandorinellina insita (Stauffer), lateral view, BT II, sample 12, IRScNB b5230. 24, 25, Ancyrodella cf. A. pristina Khalymbadzha and Chernysheva, upper and lower views, BT III, sample 2, IRScNB b5239. 26, 27, Ancyrodella pramosica Perri and Spalletta, upper and lower views, BT III, sample 8, IRScNB b5240.

Figure 16 —All elements are from Bouia Fm. of Achguig Group, from Bou Tchr...

Published: 01 July 2010
, 12 , Icriodus tafilaltensis n. sp., upper, lateral and lower views, BT II, sample 12, IRScNB b5235. 13 , 14 , Polygnathus pollocki Druce, upper and lateral views, BT II, sample 10, IRScNB b5236. 15 , Palmatolepis transitans Müller, upper view, BT II, sample 14, IRScNB b5237. 16 , 17 , 18