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Journal Article

THE UPPER GIVETIAN (MIDDLE DEVONIAN) SUBTERMINUS CONODONT ZONE IN NORTH AMERICA, EUROPE AND NORTH AFRICA

KATARZYNA NARKIEWICZ, PIERRE BULTYNCK
Published: 01 July 2010
Journal of Paleontology (2010) 84 (4): 588–625.
DOI: https://doi.org/10.1666/10-029.1
.... The holotype of Icriodus subterminus from the North Liberty beds in Iowa is most likely a specimen that was reworked from the Cedar Valley Limestone. Icriodus cedarensis and Icriodus tafilaltensis are described as new species, and the diagnoses of Icriodus excavatus and Icriodus expansus are amended. Between...
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Figure15—All elements are from Cambresèque Mb. of the Beaulieu Fm.; BN samples are from the Banc Noir Quarry, others are from the Caffiers/Ferques section, Boulonnais (France); except Fig. 6 all figures are P1 elements; scale bars are 100 µm. 1, 2, Icriodus tafilaltensis n. sp., upper and lateral views, Cédule Limestone, sample BN1, IRScNB b5214. 3–6, Ancyrodella pristina Khalymbadzha and Chernysheva, Cédule Lmst., sample BN1: 3, lower view, IRScNB b5215; 4, 5, upper and lower views, IRScNB b5216; 6, lateral view of P2 element, IRScNB b5217. 7, 8, Ancyrodella binodosa Uyeno, upper and lower views, Cédule Lmst., sample BN3 IRScNB b5218. 9, 10, 13–16, Icriodus cedarensis n. sp.: 9, 10, upper and lateral views, Cédule Lmst., sample BN1, IRScNB b5219; 13, 14, upper and lateral views, Cédule Lmst., sample BN3, IRScNB b5220; 15, 16, upper and lateral views, Cambresèque Mb., sample 32, IRScNB B5221. 11, 12, Icriodus symmetricus Branson and Mehl, upper and lateral views, Cédule Lmst., sample BN1, IRScNB b5222. 17, 18, Icriodus excavatus Weddige, upper and lateral views, Cambresèque Mb., sample 32, IRScNB b5223. 19, 20, Icriodus subterminus Youngquist α morphotype, upper (reillustration from Brice et al., 1979a, pl. 27, fig. 26, IRScNB b1132) and lateral views, Cambresèque Mb., sample 32, IRScNB b1132. 21, 22, Icriodus subterminus Youngquist, β morphotype, upper and lateral views, Cambresèque Mb., sample 32, IRScNB b5224. 23, 24, Icriodus aff. I. cedarensis, upper and lateral views, Cambresèque Mb., sample 32, IRScNB b5225.

Figure 15 —All elements are from Cambresèque Mb. of the Beaulieu Fm.; BN ...

Published: 01 July 2010
Uyeno, upper and lower views, Cédule Lmst., sample BN3 IRScNB b5218. 9 , 10 , 13–16 , Icriodus cedarensis n. sp.: 9 , 10 , upper and lateral views, Cédule Lmst., sample BN1, IRScNB b5219; 13 , 14 , upper and lateral views, Cédule Lmst., sample BN3, IRScNB b5220; 15 , 16 , upper and lateral
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Figure7—All elements from Rapid Mb., Conklin Quarry, Iowa (USA): all figures are P1 elements; scale bars are 100 µm. 1, 2, 3, Icriodus subterminus Youngquist, α morphotype, upper, lateral and lower views, SUI 100607 117328; 4–13, Icriodus subterminus Youngquist, β morphotype; 4, 5, 6, upper, lateral and lower views, SUI 100609 117337; 7, 8, 9, upper, lateral and lower views, SUI 100608 117332; 10, 11, upper and lateral views, SUI 100607 117329; 12, 13, upper and lateral views, SUI 100608 117333; 14–15, 21–26, Icriodus cedarensis n. sp.; 14, 15, upper and lateral views of paratype, SUI 100608 117334; 21, 22, 23, upper, lateral and lower views of holotype, SUI 100609 117338; 24, 25, 26, upper lateral and lower views, SUI 100607 117330; 16–17, Icriodus aff. I. subterminus Youngquist, upper and lateral views, SUI 100608 117335; 18, 19, 20, Icriodus excavatus Weddige, upper, lateral and lower views, SUI 100608 117336; 27, Icriodus tafilaltensis n. sp. upper view, SUI 100607 117331.

Figure 7 —All elements from Rapid Mb., Conklin Quarry, Iowa (USA): all fig...

Published: 01 July 2010
, 5 , 6 , upper, lateral and lower views, SUI 100609 117337; 7 , 8 , 9 , upper, lateral and lower views, SUI 100608 117332; 10 , 11 , upper and lateral views, SUI 100607 117329; 12 , 13 , upper and lateral views, SUI 100608 117333; 14 – 15 , 21–26 , Icriodus cedarensis n. sp.; 14 , 15
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Figure8—Upper and lateral views of representative specimens of morphotypes of Icriodus subterminus Youngquist and Icriodus cedarensis n. sp. from the different studied areas. Iowa—see Fig. 7.1–7.2, 7.4–7.5, 7.21–7.22; Boulonnais—See Fig. 15.19–15.20, Fig. 13.7–13.8, Fig. 15.15–15.16; Ardenne—see Fig. 11.7–11.8, Fig. 12.7–12.8, Fig. 11.5–11.6; Poland: Holy Cross Mts., Sowie Górki section, sample 4, GIUS-4-559b-SG 4/1; Elevated East European Platform, Krowie Bagno IG 1 well, depth 1333.0 m, MUZ PIG 1727.II.4, Lublin Graben, Komarów IG 1, depth 2353.5 m, MUZ PIG 1727.II.5 (see Narkiewicz and Bultynck, 2007 respectively fig. 10T, 10V); Tafilalt—see Figs. 16.4–16.5, 16.6–16.7, Figs. 17.22–17.23.

Figure 8 —Upper and lateral views of representative specimens of morphotyp...

Published: 01 July 2010
Figure 8 —Upper and lateral views of representative specimens of morphotypes of Icriodus subterminus Youngquist and Icriodus cedarensis n. sp. from the different studied areas. Iowa—see Fig. 7.1–7.2, 7.4–7.5, 7.21–7.22 ; Boulonnais—See Fig. 15.19–15.20 , Fig. 13.7–13.8 , Fig. 15.15–15.16
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Figure19—All elements are from Bou Dîb Fm. of the Ma'der area (E Anti-Atlas, Morocco); all figures are P1 elements; scale bars are 100 µm. 1, 2, Schmidtognathus wittekindti Ziegler, upper and lateral views, Bou Dîb section, Upper Member, sample 6, IRScNB b5271. 3, Schmidtognathus peracutus (Bryant), upper view, Bou Dîb section, Upper Mb., sample 5, IRScNB b5272. 4, 5, Icriodus cedarensis n. sp., upper and lateral views, Bou Dîb section, Upper Mb., sample 6, IRScNB b5273. 6, Klapperina disparalvea (Orr and Klapper), upper view, Hillock 760-Bou Terga section, Upper Mb., sample 12, IRScNB b5274. 7, 8, Polygnathus cristatus ectypus Huddle, upper and lower views, Hillock 760–Bou Terga section, Middle Mb., sample 7, IRScNB b5275. 9, 10, Polygnathus ovatinodosus, Ziegler and Klapper, upper and lateral views, Hillock 760–Bou Terga section, Middle Mb., sample 7, IRScNB b5276. 11, 12, Icriodus tafilaltensis n. sp., upper and lower views, Hillock 760–Bou Terga section, Upper Mb., sample 12, IRScNB b5277. 13, Icriodus difficilis Ziegler and Klapper, upper view, Hillock 760–Bou Terga section, Upper Mb., sample 12, IRScNB b5278. 14, 15, Polygnathus pollocki Druce, upper and lower views, Hillock 760-Bou Terga section, Upper Mb., sample 15, IRScNB b5279.

Figure 19 —All elements are from Bou Dîb Fm. of the Ma'der area (E Anti-A...

Published: 01 July 2010
peracutus (Bryant), upper view, Bou Dîb section, Upper Mb., sample 5, IRScNB b5272. 4 , 5 , Icriodus cedarensis n. sp., upper and lateral views, Bou Dîb section, Upper Mb., sample 6, IRScNB b5273. 6 , Klapperina disparalvea (Orr and Klapper), upper view, Hillock 760-Bou Terga section, Upper Mb
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Figure17—All elements are from Bouia Fm. of Achguig Group, from Bou Tchrafine section BT I of the Tafilalt area (E Anti-Atlas, Morocco); all figures are of P1 elements; scale bars are 100 µm. 1–7, 12, 13, Icriodus tafilaltensis n. sp., sample 33b: 1, 2, 3, upper, lateral and lower views of holotype, IRScNB b5241; 4, 5 and 6, 7, upper and lateral views of paratypes IRScNB b5242 and b5243, 12, 13, upper and lower views of paratype IRScNB b5244. 8, Ozarkodina semialternans (Wirth), lateral view, sample 31b, IRScNB b5245. 9, 10, Icriodus expansus Branson and Mehl, upper and lateral views, sample 33b, IRScNB b5246. 11, Icriodus difficilis Ziegler and Klapper, upper view, sample 43, IRScNB b5247. 14, 15, Polygnathus cristatus ectypus Huddle, upper and lower views, sample 35, IRScNB b5248. 16, 17, Polygnathus dengleri dengleri Bischoff and Ziegler, lateral and upper views, sample 40, IRScNB b5249. 18, Schmidtognathus wittekindti Ziegler, upper view, sample 41, IRScNB b5250. 19, Schmidtognathus latifossatus Wirth, upper view, sample 33b, IRScNB b5251. 20, 21, Icriodus subterminus Youngquist, β morphotype, upper and lateral views (reillustration from Bultynck and Hollard, 1980, pl. 8, fig. 22, IRScNB b1011), sample 37. 22, 23, Icriodus cedarensis n. sp., upper and lateral views (reillustration from Bultynck, 2003, pl. 3, fig. 22, IRScNB b4237), sample 37. 24, 25, Ancyrodella cf. A. pristina Khalymbadzha and Chernysheva, upper and lower views, sample 37, IRScNB b5252. 26, 27, Pandorinellina insita (Stauffer), upper and lateral views, sample 43, IRScNB b5253.

Figure 17 —All elements are from Bouia Fm. of Achguig Group, from Bou Tchr...

Published: 01 July 2010
views (reillustration from Bultynck and Hollard, 1980 , pl. 8, fig. 22, IRScNB b1011), sample 37. 22 , 23 , Icriodus cedarensis n. sp., upper and lateral views (reillustration from Bultynck, 2003 , pl. 3, fig. 22, IRScNB b4237), sample 37. 24 , 25 , Ancyrodella cf. A. pristina Khalymbadzha
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Figure11—All elements are from Fromelennes Fm. of the Moulin Boreux section in the Givet-Fromelennes area, Ardenne (France): all figures are P1 elements; scale bars are 100 µm. 1, 2, 21, 22, Icriodus cf. I. subterminus Youngquist: 1, 2 upper (reillustration of Gouwy, 2004, p. 124, pl. 1, fig. 1) and lateral views, Moulin Boreux Mb., sample MB-ben, IRScNB b5178; 21, 22, upper (reillustration of Gouwy, 2004, p. 124, pl. 1, fig. 5) and lateral views, Fort Hulobiet Mb., sample MB2-5, IRScNB b5179. 3, 4, 9, 10, 15, 16, Icriodus subterminus Youngquist, β morphotype: 3, 4, upper and lateral views, Moulin Boreux Mb., sample MB PB1, IRScNB b5180; 9, 10, upper and lateral views, Fort Hulobiet Mb., sample MBH, IRScNB b5181; 15, 16, upper and lateral views, Fort Hulobiet Mb., sample MB PB2, IRScNB b5182. 5, 6, 11, 12, 17, 18, Icriodus cedarensis n. sp.: 5, 6, upper and lower views, Moulin Boreux Mb., sample MB-ben, IRScNB b5183; 11, 12, upper and lateral views, Fort Hulobiet Mb., sample MB2-5, IRScNB b5184; 17, 18, upper and lateral views, Fort Hulobiet Mb., sample MB PB2, IRScNB b5185. 7, 8, 13, 14, Icriodus subterminus Youngquist, α morphotype from Fort Hulobiet Mb.: 7, 8, upper and lateral views, sample MBH, IRScNB b5186; 13, 14, sample MB PB2, IRScNB b5187. 19, 20, Ozarkodina brevis (Bischoff and Ziegler), upper and lateral views, Fort Hulobiet Mb., sample MB PB2, IRScNB b5188. 23, 24, Polygnathus pollocki Druce from Fort Hulobiet Mb.: upper view (Fig. 24, reillustration of Gouwy, 2004, p. 124, pl. 1, fig. 10), sample MB2-5, IRScNB b5189, IRScNB b5190.

Figure 11 —All elements are from Fromelennes Fm. of the Moulin Boreux sect...

Published: 01 July 2010
cedarensis n. sp.: 5 , 6 , upper and lower views, Moulin Boreux Mb., sample MB-ben, IRScNB b5183; 11 , 12 , upper and lateral views, Fort Hulobiet Mb., sample MB2-5, IRScNB b5184; 17 , 18 , upper and lateral views, Fort Hulobiet Mb., sample MB PB2, IRScNB b5185. 7 , 8 , 13 , 14 , Icriodus
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Figure14—All figures are P1 elements; scale bars are 100 µm. 1–12, 18–23, Icriodus lilliputensis Bultynck: 1, 2, 3, upper, lateral and lower views, Hotton/Ny section, Fort Hulobiet Mb. of the Fromelennes Fm., sample 27, Ardenne (Belgium), IRScNB b5264; 4, upper view (lateral view in Bultynck, 1987, pl. 5, fig. 19), Mont d'Haurs section, Mont d'Haurs Fm., sample 43, Ardenne (France), IRScNB b1898; 5, 6, upper and lateral views, Hotton/Ny section, Fort Hulobiet Mb. of the Fromelennes Fm., sample 27, Ardenne (Belgium), IRScNB b5265; 7–10, are from section 39, Micrit Limestone Beds of the Point Wilkins Mm. of the Souris River Fm., Manitoba (Canada): 7, 8, upper and lateral views, sample GSC loc. 78896, 0.91–1.92 m above base of the section, IRScNB b5266; 9, 10, upper and lateral views, sample GSC loc. 78897, 1.92–2.83 m above base of the section, IRScNB b5267; 11, 12, upper and lateral views, Birch River area, Unit IV of the Waterways Fm., sample GSC 74172, NE Alberta (Canada), IRScNB b5268; 18, 19, upper and lateral views, Krowie Bagno IG 1 well, Telatyn Fm., depth 1332.2 m, Elevated East European Platform (Poland), MUZ PIG 1727.II.2; 20, 21, upper (reillustration of Narkiewicz and Bultynck, 2007, fig. 10D) and lateral views, Krowie Bagno IG 1 well, Telatyn Fm., depth 1333.0 m, Elevated East European Platform (Poland), MUZ PIG 1727.II.1; 22, 23, upper and lateral views, Giełczew PIG 5 well, Telatyn Fm., depth 1967.1 m, Lublin Graben (Poland), MUZ PIG 1727.II.3. 13, 14, 15, Icriodus excavatus Weddige, upper, lateral and lower views, Unit II of the Waterways Fm., sample GSC loc. 74175, 9.4–9.6 m above base of the section, Lake Clair and Birch River area, NE Alberta (Canada), IRScNB b5269. 16, 17, Icriodus cedarensis n. sp., upper and lateral views, section 39, Micrit Limestone Beds of the Point Wilkins Mb of Souris River Fm., sample GSC 78898, Manitoba (Canada), IRScNB b5270. 24, Icriodus subterminus Youngquist, upper view, Komarów IG 1, Telatyn Fm., depth 2353.5 m, Lublin Graben (Poland), MUZ PIG 1727.II.6.

Figure 14 —All figures are P 1 elements; scale bars are 100 µm. 1–12 , ...

Published: 01 July 2010
, 14 , 15 , Icriodus excavatus Weddige, upper, lateral and lower views, Unit II of the Waterways Fm., sample GSC loc. 74175, 9.4–9.6 m above base of the section, Lake Clair and Birch River area, NE Alberta (Canada), IRScNB b5269. 16 , 17 , Icriodus cedarensis n. sp., upper and lateral views