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Cissites puilasokensis

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Late Campanian–early Maastrichtian paleogeography of the Western Interior Seaway (WIS; modified from Slattery et al., 2015). Key to numbered records of “Cissites” puilasokensis or “Cissites” panduratus: (1)Heer (1883): early Paleocene; (2)Knowlton (1917), Lee (1917), Wolfe and Upchurch (1987): late Maastrichtian–early Paleocene; (3)Knowlton (1917), Lee (1917), Wolfe and Tanai (1987): early Paleocene; (4)Berry (1922, 1925): late Campanian or early Maastrichtian; (5)Brown (1939): late Maastrichtian; (6)Barclay et al. (2003), Nichols and Johnson (2008): early Paleocene; (7)Johnson and Hickey (1990), Nichols and Johnson (2008): late Maastrichtian. When reviewing these records, which are arranged more or less chronologically, please note that Berry (1922, p. 171) erroneously reported that “C.” panduratus initially had been described from the Mesaverde Formation (no locality given), but that he later corrected this report by acknowledging that, at the time, “C.” panduratus was known only from the Vermejo and Ripley Formations and not the Mesaverde Formation (Berry, 1925, p. 24). Key to lettered paleogeographic features: (a) northern shoreline of the WIS according to Erickson (1978, 1999), Lillegraven and Ostresh (1990), Robinson Roberts and Kirschbaum (1995), and Kennedy et al. (1998); (b) southern shoreline of the WIS according to Lehman (1987) and Blakey (2014); (c) dispersal route for land plants between Asia and western North America in the late Maastrichtian (Peppe et al., 2007; Zakharov et al., 2011); (d) eastern margin of the WIS and general location (e) of a large island or archipelago according to Williams and Stelck (1975). Local paleogeographic map (modified from Miller and McKinney, 2016): Position of western shoreline of the WIS during the Baculites clinolobatus ammonite biozone as reconstructed by Cobban et al. (1994), Landman and Cobban (2003), Slattery et al. (2015), and Miller and McKinney (2016). New data from the Raton Basin, which are discussed in the text, indicate that this shoreline should be shifted about 100 km farther west (white arrow) to a geographic location in the immediate vicinity of Berwind Canyon, Colorado.
Published: 01 June 2017
Figure 1. Late Campanian–early Maastrichtian paleogeography of the Western Interior Seaway (WIS; modified from Slattery et al., 2015 ). Key to numbered records of “Cissitespuilasokensis or “Cissites” panduratus : (1) Heer (1883) : early Paleocene; (2) Knowlton (1917) , Lee (1917
Journal Article
Published: 01 June 2017
Rocky Mountain Geology (2017) 52 (1): 1–16.
...Figure 1. Late Campanian–early Maastrichtian paleogeography of the Western Interior Seaway (WIS; modified from Slattery et al., 2015 ). Key to numbered records of “Cissitespuilasokensis or “Cissites” panduratus : (1) Heer (1883) : early Paleocene; (2) Knowlton (1917) , Lee (1917...
FIGURES
First thumbnail for: New paleontological constraints on the paleogeogra...
Second thumbnail for: New paleontological constraints on the paleogeogra...
Third thumbnail for: New paleontological constraints on the paleogeogra...
Image
A biostratigraphically important leaf fossil from the Vermejo-Raton megafloras. A, “Cissites” panduratus (NMMNH P-78592) from the upper coal zone (Paleocene) of the Raton Formation in Road Canyon, Colorado, which merges with Berwind Canyon to the east (see Fig. 1; NMMNH Locality L-10794). Weakly lobed leaves, such as this one, from the upper coal zone of the Raton Formation in south-central Colorado, originally had been referred to “Acer” fragilis by Knowlton (1917) and Lee (1917); however, this taxon was later synonymized with “C.” panduratus by Wolfe and Tanai (1987)—a taxonomic revision that still stands (Manchester, 2014). B, Original figures of the deeply lobed type (left) and plesiotype (right) specimens of “C.” panduratus from the Vermejo and Ripley Formations, respectively (reproduced from Knowlton, 1917, plate 69, fig. 10; Berry, 1925, plate 14, figs. 2–3). C, Original figure of the type specimen of “C.” puilasokensis from the Puilasok megaflora (Paleocene) of western Greenland (reproduced from Heer, 1883, plate 107, fig. 8), which is now known to be early Paleocene in age (see LePage et al., 2005). In the Raton Basin, deeply lobed leaves essentially identical to this morphotype have been attributed to “C.” panduratus and are locally abundant in Wolfe and Upchurch’s (1987) early Paleocene megafloral assemblage in the Raton Formation (Upchurch, personal communication, 2014). Consequently, “C.” puilasokensis and Wolfe and Upchurch’s (1987) early Paleocene assemblage, which is dominated by “C.” panduratus, are now known to be essentially equivalent in age (Fig. 1).
Published: 01 June 2017
Figure 6. A biostratigraphically important leaf fossil from the Vermejo-Raton megafloras. A , “Cissites” panduratus (NMMNH P-78592) from the upper coal zone (Paleocene) of the Raton Formation in Road Canyon, Colorado, which merges with Berwind Canyon to the east (see Fig. 1 ; NMMNH Locality