... sutural surface to the tip of the distally widening scapular blade (RSKM P3217.560, Figs. 10A – 10C ). A slightly rounded distal end to the scapula suggests a closer taxonomic affinity to Centrosaurinae than to Chasmosaurinae. The large size of the specimen combined with the absence of a coracoid fused...

...Michael J. Ryan; Robert Holmes; Jordan Mallon; Mark Loewen; David C. Evans Abstract A partial skull (CMN 8804) of a ceratopsid from the upper unit of the Campanian Oldman Formation of Alberta is the first Canadian example of the newly established Nasutoceratopsini , a new subclade of Centrosaurinae...

... of north-central Montana. A limited phylogenetic analysis of the Ceratopsidae places the new taxon as the basal member of the Centrosaurinae and indicates that robust, elongate postorbital horncores that form a synapomorphy of (Ceratopsidae + Zuniceratops ) are also present in Centrosaurinae...

... that are characteristic of Centrosaurinae. From medial to lateral, the epiparietal ornamentation consists of a small, variably procurving epiparietal (ep 1), an anterolaterally curving pachyostotic hook (ep 2), a smaller pachyostoic process (ep 3), and two small triangular epiparietals (ep 4 and 5). A phylogenetic...

...Fig. 7. Reconstructions of Centrosaurinae parietals. (A) Xenoceratops foremostensis ; (B) Centrosaurus apertus ; (C) Styracosaurus albertensis ; (D) Achelousaurus horneri ; (E) Albertaceratops nesmoi ; (F) Pachyrhinosaurus lakustai ; (G) Einiosaurus procurvicornus ; (H) Diabloceratops...

...Fig. 10. Cladistic relationships of the Centrosaurinae. Characters (1–17) and states (superscripts 0–4) as outlined in Appendix A. Percent support for clades from bootstrap analysis (5000 replicates) are listed in italics above each of the branches. ...

... that in Centrosaurinae and cannot be used as a tool for distinguishing the two ceratopsid subfamilies. Nous rapportons la première découverte dans la Formation de Frenchman (Maastrichtien tardif) de cératppsiens juvéniles (Chasmosaurinés) identifiés sur la base de deux noyaux de cornes supraorbitaires. Les sutures...

... epiossifications ( Sampson 1995 ). Ceratopsidae consists of the subfamilies Centrosaurinae and Chasmosaurinae, which can be differentiated in having a relatively short, highly ornamented frill, and a relatively long, lesser ornamented frill, respectively ( Ryan et al. 2010 ). The Upper Cretaceous (Campanian...

...Fig. 4. Time-calibrated phylogeny of Chasmosaurinae from Sampson et al. (2010) . Figure reproduced by permission of Scott Sampson. A similar figure for Centrosaurinae appears in Sampson and Loewen (2010) . ...

... of North America's fauna. Ceratopsids are particularly diverse in the late Campanian (Judithian) of Western Canada. The short-frilled Centrosaurinae dominate in terms of numbers, but the long-frilled Chasmosaurinae are a conspicuous part of this fauna as well ( Currie and Russell, 2005 ). Currently, four...

... ). These sinuses are relatively small, simple chambers in most members of the two ceratopsid clades Chasmosaurinae and Centrosaurinae ( Lehman, 1990 ). However, the frontal sinuses reach large proportions in some species, with a portion of the sinuses extending into the base of the postorbital horncores ( Forster...

... Centrosaurinae and Chasmosaurinae, underwent a period of rapid evolution during the late Campanian and early Maastrichtian. Both centrosaurine and chasmosaurine ceratopsids were relatively common at that time, including the centrosaurines Achelousaurus, Avaceratops, Centrosaurus, Pachyrhinosaurus...

... level pertaining to Centrosaurinae ( Table 1 ), and two species-level diagnostic parietals (TMP 2012.037.0021 and TMP 2012.037.0197, Fig. 4 ) are referable to C. apertus (see Taxonomic Identification of McPheeters Bonebed Specimens). Based on the minimum estimated areal extent of the bonebed...

... Brachyceratops and Monoclonius ) distributed between the subfamilies Centrosaurinae and Chasmosaurinae. The centrosaurines have figured prominently throughout the history of North American dinosaur work, from the description of the first horned dinosaur Monoclonius ( Cope 1876 ; = Centrosaurus ) to the most...

.... Ceratopsids are large-bodied, ornithischian herbivores traditionally divided into two monophyletic clades: the ‘long-frilled’ Chasmosaurinae ( Lambe, 1915 ), and the ‘short-frilled’ Centrosaurinae ( Lehman, 1990 ). The Chasmosaurinae includes at least nine species distributed among seven genera...