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Beothuka

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Journal Article
Published: 01 November 2005
Geological Magazine (2005) 142 (6): 711–721.
...JORG MALETZ; DAVID L. BRUTON Abstract The radiolarian Beothuka terranova occurs in the Arenigian Didymograptellus bifidus Biozone (uppermost Lower Ordovician) of Spitsbergen (Svalbard), associated with a diverse and well-preserved radiolarian fauna. The presence of typical Cambrian spicular...
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Figure 3. The relation between beothukid and antygoporid shell wall meshwork. The beothukid shell wall construction is a mix of three-dimensional network with bandages and a shell wall with wide interpore areas that is the result of coalescence of “bandages” into a thin layer. Where the interpore area is very wide compared to pore size, it appears as perforate plate wall. Depending on whether the mesh with bandages or a latticed wall with wide interpore areas is dominant, the shell wall meshwork structure appears diverse. Where only perforated plate shell is developed, the antygoporid shell wall structure appears. Scale bar is 50 µm. 1a, b, Beothuka terranova terranova s.l. Aitchison et al., 1988, not well preserved, but bandage mesh is dominant, wide interpore area (dotted area in 1b); 2a, b, Beothuka sp., a shell wall with wide interpore areas dominant (dotted area in 2b); 3, Beothuka aitchisoni n. sp., incompletely layered beothukid shell structure; 4, Beothuka stougei n. sp., perforate plate wall dominant, but not completely layered to form multiple shells; 5, 6, Antygopora sp., completely layered antygoporid shell wall structure, only a shell wall with wide interpore areas developed.
Published: 01 January 2011
wall structure appears. Scale bar is 50 µm. 1a, b, Beothuka terranova terranova s.l. Aitchison et al., 1988, not well preserved, but bandage mesh is dominant, wide interpore area (dotted area in 1b); 2a, b, Beothuka sp., a shell wall with wide interpore areas dominant (dotted area in 2b); 3
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Figure 8. Scale bar equals 50 µm. 1a, b–5, 9, Beothuka terranova terranova s.l. Aitchison et al., 1998: 1a, b, two-dimensional bandage mesh in the dotted circle, GSC 131998; 2a, b, bandage (arrow) where mesh is three-dimensional and here mesh has wide interpore areas in two-dimensional structure (dotted circle), GSC 131999; 3a, b, c, d, typical bandage mesh (in 3b), some inner spines (arrow in 3b and arrows in 3c) are not developed as outer spines, and the innermost part of the shell wall consists of thin bars (arrows in 4d) and bandage, a bar may occupy the middle of the bandage (see dotted circle in 3d), GSC 131000; 4a, b, one main and two minor spines, GSC 131001; 5, one main and five minor spines, GSC 131002; 9, thorny surface structure is similar to that of B. t. multispinosa in Fig. 8.7 and 8.8, GSC 131003; 6–8, Beothuka terranova multispinosa n. ssp.; 6, Meshwork structure similar to that of Fig. 8.2a, GSC 131004; 7, 8, rough and thorny surface similar to that of Fig. 8.9; 10–12, ?16a, b, Beothuka stougei n. sp.; 10, GSC 131005; 11, Holotype, GSC 131006; 12, GSC 131007; 16a, b, note the connecting element between layers; the original bandage shape of the connecting element is modified to trapezoidal (black arrow) or to a thin bar near the margin of the pore (white arrow), GSC 131008; 13–15, Beothuka aitchisoni n. sp.; 13, GSC 1310009; 14, holotype, GSC 131010; 15, GSC 131012.
Published: 01 January 2011
Figure 8. Scale bar equals 50 µm. 1a, b–5, 9, Beothuka terranova terranova s.l. Aitchison et al., 1998 : 1a, b, two-dimensional bandage mesh in the dotted circle, GSC 131998; 2a, b, bandage (arrow) where mesh is three-dimensional and here mesh has wide interpore areas in two-dimensional
Journal Article
Published: 01 January 2011
Journal of Paleontology (2011) 85 (1): 156–177.
... wall structure appears. Scale bar is 50 µm. 1a, b, Beothuka terranova terranova s.l. Aitchison et al., 1988, not well preserved, but bandage mesh is dominant, wide interpore area (dotted area in 1b); 2a, b, Beothuka sp., a shell wall with wide interpore areas dominant (dotted area in 2b); 3...
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Figure 9. Scale bar equals 50 µm. 1–7a, b, Beothuka grosmornensis n. sp.: 1, GSC 131013; 2, GSC 131014; 3, GSC 131015; 4, GSC 131016; 5, holotype, GSC 131017; 6, GSC 131018; 7a, b, mesh with thin bar, narrow bandage, and wide interpore area, GSC 131019; 8–12a, b, Beothuka? stellata n. sp.; 8, holotype, GSC 131020; 9, GSC 131021; 10, GSC 131022; 11, GSC131023; 12a, b, proloculus constructed by thin bars, GSC 131024; 13, 14–16a, b, Neoaspiculum cancellium n. sp.; 13, GSC131025; 14, GSC 131026; 15a, b, enlarged meshwork of the inner shell shows incomplete two-dimensional subspherical shape; holotype, GSC 131027; 16a, b, eccentric inner shell, GSC 131028.
Published: 01 January 2011
Figure 9. Scale bar equals 50 µm. 1–7a, b, Beothuka grosmornensis n. sp.: 1, GSC 131013; 2, GSC 131014; 3, GSC 131015; 4, GSC 131016; 5, holotype, GSC 131017; 6, GSC 131018; 7a, b, mesh with thin bar, narrow bandage, and wide interpore area, GSC 131019; 8–12a, b, Beothuka
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Lithostratigraphy of the Vahallfonna Formation (based on Fortey & Bruton, 1973) with graptolite biostratigraphy revised from Cooper & Fortey (1982). 1 – Beothuka terranova radiolarian level. 2 – Isograptus victoriae lunatus Biozone radiolarians, not described herein. 3 – Proventocitum procerulum radiolarian fauna described by Fortey & Holdsworth (1971).
Published: 01 November 2005
Figure 1. Lithostratigraphy of the Vahallfonna Formation (based on Fortey & Bruton, 1973 ) with graptolite biostratigraphy revised from Cooper & Fortey (1982) . 1 – Beothuka terranova radiolarian level. 2 – Isograptus victoriae lunatus Biozone radiolarians, not described herein. 3
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Lower Paleozoic radiolarian assemblages of Gorny Altai and their skeleton preservation. A, Shashkunar Formation: figs. 1, 4, Archaeocenosphaera muricata Obut et Iwata; fig. 2. Altaiesphaera sparsispinosa Obut et Iwata; fig. 3, Altaiesphaera acanthophora Obut et Iwata; fig. 5, 6, Parechindina sp. Scale bar 50 µm; B, Zasur’ya Group: figs. 1, 2, Praespongocoelia sp.; fig. 3, Beothuka sp.; fig. 4, Inanigutta sp.; scale bar 100 µm; C, Tekhten’ Formation and the siliceous–terrigenous strata: figs. 1, 2, Secuicollacta ornata Goto, Umeda et Ishiga; figs. 3, 5, Kalimnasphaera cf. maculosa Web. et Bl.; fig. 4, Protoceratoikiscum chinocrystallum Goto, Umeda et Ishiga; figs. 6, 7, Borisella subulata (Web. et Bl.); scale bar 100 µm.
Published: 01 October 2011
. 5 , 6, Parechindina sp. Scale bar 50 µm; B , Zasur’ya Group: figs. 1 , 2 , Praespongocoelia sp.; fig. 3 , Beothuka sp.; fig. 4 , Inanigutta sp.; scale bar 100 µm; C , Tekhten’ Formation and the siliceous–terrigenous strata: figs. 1 , 2 , Secuicollacta ornata Goto, Umeda et Ishiga
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Upper Cambrian to lower Ordovician radiolarian faunas and time scale. For the highly diverse Cambrian faunas of Won & Iams (2002) only the genus names are given for easier comparison. Grey underlain areas show uncertainties in biostratigraphic ranges. Dotted lines indicate known ranges of genera. Specimens: (a) Antygopora sp., JM 3/60, western Newfoundland, Isograptus victoriae maximodivergens Biozone. (b) Antygopora ordovicica n. sp., JM 3/68, western Newfoundland, Isograptus victoriae maximodivergens Biozone. (c) Secuicollacta sp. Llandovery, Dalarna, Sweden. (d) Entactinid species indet., PMO 205.716/44, Spitsbergen, 102 m level. (e) Beothuka terranova, PMO 205.718/58, Spitsbergen. (f ) Palaeospiculum sp. PMO 205.715/55, Spitsbergen, 102 m level. (g) Subechidnina florea, western Newfoundland (Won & Iams, 2002).
Published: 01 November 2005
) Entactinid species indet., PMO 205.716/44, Spitsbergen, 102 m level. (e) Beothuka terranova , PMO 205.718/58, Spitsbergen. (f ) Palaeospiculum sp. PMO 205.715/55, Spitsbergen, 102 m level. (g) Subechidnina florea , western Newfoundland ( Won & Iams, 2002 ).
Journal Article
Published: 01 July 2003
Bulletin de la Société Géologique de France (2003) 174 (4): 325–335.
... N azarov . L’absence du genre Beothuka – un nouveau genre récemment décrit du Trémadocien inférieur [Aitchison et al. 1998] – et des Inaniguttidae à six épines primaires ( Inanigutta , Inanibigutta ) est remarquable. L’assemblage qui suit dans le temps correspond à celui définit par...
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(a–d) Antygopora ordovicica n. gen. n. sp.: (a) PMO 205.718/13, holotype, showing two tests connected by few beams. (b) PMO 205.718/22, outer test with honeycomb surface structure. (c) PMO 205.718/44, specimen with broken outer sphere. (d) PMO 205.717/00, surface of inner sphere with several beams. (e, f ) Beothuka terranovaAitchison, Flood & Malpas, 1998: (e) PMO 205.718/58, typical example. (f ) PMO 205.718/129, broken specimen showing inside and grooved base of spine. (g) Antygopora (?) sp. PMO 205.718/73, specimen with one test bearing large pores with raised rims and about 10–12 spines. Indication of inner beams or spicule is visible on lower left side. (h, j) Parechidnina sp. B: (h) PMO 205.717/188, complete specimen with short spines. (j) PMO 205.717/08, juvenile form? (i) Parechidnina sp. A, PMO 205.718/25, long spine with labyrinthine mesh at base. (k) Palaeospiculum sp., PMO 205.718/28. (l) ?Entactinia sp., PMO 205.718/190, showing ectopically placed spicule with six rays. (m) ?Parechidnina sp., PMO 205.718/107, fragment. (n) Ramuspiculum sp., PMO 205.718/59. (o) PMO 205.717/43, re-crystallized sphere indet.
Published: 01 November 2005
with several beams. (e, f ) Beothuka terranova Aitchison, Flood & Malpas, 1998 : (e) PMO 205.718/58, typical example. (f ) PMO 205.718/129, broken specimen showing inside and grooved base of spine. (g) Antygopora (?) sp. PMO 205.718/73, specimen with one test bearing large pores with raised rims
Journal Article
Published: 01 November 2008
Journal of Micropalaeontology (2008) 27 (2): 97–101.
.... & Bruton , D.L. 2005 . The Beothuka terranova (Radiolaria) assemblage and its importance for the understanding of early Ordovician radiolarian evolution . Geological Magazine , 142 : 1 – 11 . Overstreet , R.B. , Obuh-Ikuenobe , F.E. & Gregg , J.M. 2003 . Sequence...
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Journal Article
Published: 01 September 2021
Journal of Paleontology (2021) 95 (5): 913–943.
... (Radiolaria) from the Lower Silurian of the Cape Phillips Formation, Cornwallis Island, Nunavut, Canada : Journal of Paleontology , v. 80 , p. 19 – 37 . Maletz , J. , 2007 , The Early Ordovician Beothuka terranova (Radiolaria) faunal assemblage in Newfoundland : Palaeontologische Zeitschrift...
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Journal Article
Published: 01 January 2022
Journal of Paleontology (2022) 96 (1): 19–45.
... (Radiolaria) from the lower Silurian of the Cape Phillips Formation, Cornwallis Island, Nunavut, Canada : Journal of Paleontology , v. 78 , p. 257 – 274 . Maletz , J. , 2007 , The Early Ordovician Beothuka terranova (Radiolaria) faunal assemblage in western Newfoundland : Palaeontologische...
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Journal Article
Published: 29 April 2020
Geological Magazine (2020) 157 (12): 2033–2043.
... that the dense mesh can be readily separated into three whorls. Without the aid of a micro-CT model, it is very challenging to differentiate primary and secondary spines. This species also bears some superficial similarity with Beothuka echinata Won & Iams ( 2013 ) as a result of the incomplete layering...
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Series: Geological Society, London, Memoirs
Published: 01 January 2013
DOI: 10.1144/M38.25
EISBN: 9781862396425
... ) Parechidnina jamesi ( Won & Iams 2002 ); ( e ) Oriundogutta miscella miscella ( Nazarov, in Nazarov & Popov 1980 ); ( f ) Inanigutta gansuensis ( Wang 1993 ); ( g ) Antygopora bella ( Maletz & Bruton 2007 ); ( h ) Sphaeroentactinia hexaspinosa ( Maletz & Bruton 2007 ); ( i ) Beothuka...
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Journal Article
Published: 01 October 2011
Russ. Geol. Geophys. (2011) 52 (10): 1171–1194.
.... 5 , 6, Parechindina sp. Scale bar 50 µm; B , Zasur’ya Group: figs. 1 , 2 , Praespongocoelia sp.; fig. 3 , Beothuka sp.; fig. 4 , Inanigutta sp.; scale bar 100 µm; C , Tekhten’ Formation and the siliceous–terrigenous strata: figs. 1 , 2 , Secuicollacta ornata Goto, Umeda et Ishiga...
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Journal Article
Published: 01 May 2020
Journal of Paleontology (2020) 94 (3): 417–435.
...) from the Lower Silurian of the Cape Phillips Formation, Cornwallis Island, Nunavut, Canada : Journal of Paleontology , v. 80 , p. 19 – 37 . doi: 10.1666/0022- 3360(2006)080[0019:HAIRFT]2.0.CO;2 Maletz , J. , 2007 , The Early Ordovician Beothuka terranova (Radiolaria) faunal assemblage...
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