Abstract The most extensive Permian tetrapod (amphibian and reptile) fossil records from the western United States (New Mexico–Texas) and South Africa provide the basis for definition of 10 landvertebrate faunachrons that encompass Permian time. These are (in ascending order): the Coyotean, Seymouran, Mitchellcreekian, Redtankian, Littlecrotonian, Kapteinskraalian, Gamkan, Hoedemakeran, Steilkransian and Platbergian. These faunachrons provide a biochronological framework with which to determine and discuss the age relationships of Permian tetrapod faunas. Their correlation to the marine time scale and its numerical calibrations indicate that the Coyotean is a relatively long time interval of about 20 Ma, whereas most of the other faunachrons are much shorter, about 1–2 Ma long each. The Platbergian may also be relatively long, 14 Ma, although this is not certain. This suggests slow rates of terrestrial tetrapod faunal turnover during most of the Early Permian and late Middle to Late Permian, but more rapid rates of turnover during the latest Early and most of the Middle Permian, especially during the explosive initial diversification of therapsids.

Abstract: The most extensive Permian tetrapod (amphibian and reptile) fossil records from the western USA (New Mexico to Texas) and South Africa have been used to define 11 land vertebrate faunachrons (LVFs). These are, in ascending order, the Coyotean, Seymouran, Mitchellcreekian, Redtankian, Littlecrotonian, Kapteinskraalian, Gamkan, Hoedemakeran, Steilkransian, Platbergian and Lootsbergian. These faunachrons provide a biochronological framework with which to assign ages to, and correlate, Permian tetrapod fossil assemblages. Intercalated marine strata, radioisotopic ages and magnetostratigraphy were used to correlate the Permian LVFs to the standard global chronostratigraphic scale with varying degrees of precision. Such correlations identified the following significant events in Permian tetrapod evolution: a Coyotean chronofaunal event (end Coyotean); Redtankian events (Mitchellcreekian–Littlecrotonian); Olson’s gap (late Littlecrotonian); a therapsid event (Kapteinskraalian); a dinocephalian extinction event (end Gamkan); and a latest Permian extinction event (Platbergian–Lootsbergian boundary). Problems of incompleteness, endemism and taxonomy, and the relative lack of non-biochronological age control continue to hinder the refinement and correlation of a Permian timescale based on tetrapod biochronology. Nevertheless, the global Permian timescale based on tetrapod biochronology is a robust tool for both global and regional age assignment and correlation. Advances in Permian tetrapod biochronology will come from new fossil discoveries, more detailed biostratigraphy and additional alpha taxonomic studies based on sound evolutionary taxonomic principles.