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Azhdarchidae
A large pteranodontid pterosaur from the Late Cretaceous of Eastern Europe
The pterosaurian remains from the Grünbach Formation (Campanian, Gosau Group) of Austria: a reappraisal of ‘ Ornithocheirus buenzeli ’
An early azhdarchid pterosaur from the Lower Cretaceous of the eastern Paris basin
A wing metacarpal from Italy and its implications for latest Cretaceous pterosaur diversity
Abstract: An incomplete bone from the latest Cretaceous dinosaur site of Villaggio del Pescatore (Trieste Province, Italy) is definitely a wing metacarpal of a pterodactyloid pterosaur. It represents the only Italian Cretaceous pterosaur remains known, as well as the only pterosaur from the Adriatic Carbonate Platform. With an estimated minimum length of 136 mm, it belongs to a relatively small individual relative to the standard of latest Cretaceous pterodactyloids. It is not as elongated and gracile as azhdarchid wing metacarpals and shows a mix of features found in Pteranodon and some more basal pterodactyloids. It is one of the very few remains of putative non-azhdarchid pterosaurs from the upper Campanian–Maastrichtian worldwide and supports the view that the Azhdarchidae were not the only pterosaur clade existing during latest Cretaceous times.
Biogeographical affinities of Late Cretaceous continental tetrapods of Europe: a review
Abstract A large azhdarchid pterosaur is described from the Late Maastrichtian phosphatic deposits of the Oulad Abdoun Basin, near Khouribga (central Morocco). The material consists of five closely associated cervical vertebrae of a single individual. The mid-series neck vertebrae closely resemble those of azhdarchids Quetzalcoatlus and Azhdarcho in that they are elongate, with vestigial neural spines, prezygapophysial tubercles, a pair of ventral sulci near the prezygapophy ses, and without pneumatic foramina on the lateral surfaces of the centra. The Moroccan pterosaur is referred to a new genus and species of Azhdarchidae: Phosphatodraco mauritanicus gen. et sp.nov. It is mainly characterized by a very long cervical vertebra eight, bearing a prominent neural spine located very posteriorly. Based on comparisons with azhdarchid vertebrae, the estimated wing span of Phosphatodraco is close to 5 m. This discovery provides the first occurrence of Late Cretaceous azhdarchids in northern Africa. Phosphatodraco is one of the few azhdarchids known from a relatively complete neck and one of the latest-known pterosaurs, approximately contemporaneous with Quetzalcoatlus .
Pterosaur phylogeny and comments on the evolutionary history of the group
Abstract A cladistic analysis based on 39 terminal taxa and 74 characters (several multistate) using PAUP (Phylogenetic Analysis Using Parsimony) (3.1.1 for Macintosh and 4.0b10 for Microsoft Windows) presents a new hypothesis of pterosaur inter-relationships. This study suggests that the most primitive taxon is the Anurognathidae, followed by Sordes and all remaining pterosaurs. Dendrorhynchoides is confirmed as a member of the Anurognathidae, being closely related to Batrachognathus. Preondactylus occupies a more derived position than Sordes, which questions its previous assignment as the most primitive pterosaur. The hypothesis of rhamphorhynchoid paraphyly is confirmed, with the Rhamphorhynchidae more closely related to the Pterodactyloidea than to more basal forms. The Pterodactyloidea shows a basal dichotomy: the Archaeopterodactyloidea and the Dsungaripteroidea. The Archaeopterodactyloidea is formed by Pterodactylus + Germanodactylus and a clade formed by Gallodactylidae + Ctenochasmatidae. The Nyctosauridae occupies the basal position within dsungaripteroids and is followed by the Pteranodontoidea and the Tapejaroidea. Pteranodontoids have Pteranodon at the base, followed stepwise by Istiodactylus, Ornithocheirus and the Anhangueridae. Tapejaroids are composed of the Dsungaripteridae at the base followed by the Tapejaridae and the Azhdarchidae. Major trends within pterosaur evolutionary history are: general increase in size (wing span and body); increase of wing metacarpal and pteroid; decrease of proportional length of the second and third wing phalanx relative to the first; gradual increase of rostrum (anterior to external nares); and anterior shift of the skull-mandible articulation. Cranial crests are present in most pterodacty-loids, but markedly in the Ornithocheiroidea, where all taxa show some sort of crest on the skull. The loss of teeth, previously assumed to have occurred independently in several lineages, seems to be a general trend among dsungaripteroids. Several nodes recovered by this analysis are supported by very few characters, a result at least partially attributable to the limited available information from several taxa due to poor preservation and/or preparation.
Abstract Previous cladistic studies of pterosaur relationships suffer from restricted numbers of taxa and characters, incomplete data sets and absence of information on characters, tree structure and the robustness of trees. Parsimony analysis of a new character data set (60 characters, 20 terminal taxa, 93.75% complete) yielded six trees. In the strict consensus tree Preondactylus is the most basal taxon followed, stepwise, by the Dimorphodontidae and the Anurognathidae. Beyond this basal group, more derived pterosaurs (Campylognathoididae (Rhamphorhynchidae + Pterodactyloidea)) share a suite of characters principally associated with elongation of the rostrum. The Pterodactyloidea consists of four major clades. The Ornithocheiroidea is the most basal taxon consisting, stepwise, of Istiodactylus, the Ornithocheiridae, Nyctosaurus and the Pteranodontidae. The remaining taxa, Ctenochasmatoidea, Dsungaripteroidea and Azhdarchoidea, are weakly united in a clade of non-ornithocheiroid pterodactyloids, but their inter-relationships are difficult to resolve. Cycnorhamphus is the basal-most ctenochasmatoid, while the remaining taxa (Pterodactylus, Lonchodectidae, Ctenochasmatidae) form an unresolved trichotomy. The Dsungaripteroidea (Germanodactylus + Dsungaripteridae) is strongly supported by features of the skull and dentition. The Azhdarchoidea (Tapejara [Tupuxuara + Azhdarchidae]) is united by cranial characters such as elevation of the antorbital region, and relative shortening of the wing finger. The pattern of pterosaur evolution suggested by the results of this analysis is broadly similar to traditional ideas, but has greater resolution, more complexity and reveals several previously unrecognized ‘events’.
Fig. 6. Stratigraphic distribution of pterosaur clades through time, show...
Estimating the effects of sampling biases on pterosaur diversity patterns: implications for hypotheses of bird/pterosaur competitive replacement
Earliest Occurrence of the Pteranodontidae (Archosauria: Pterosauria) in North America: New Material from the Austin Group of Texas
Cervical vertebrae of an enigmatic pterosaur from the Crato Formation (Lower Cretaceous, Araripe Basin, NE Brazil)
Abstract: The Brazilian Crato Formation (Lower Cretaceous, Aptian) is well known for its rich pterosaur fauna. This paper deals with a new find represented by four articulated mid-cervical vertebrae. The vertebrae show a morphology consistent with that seen in the Chaoyangopteridae, especially the relative elongation, low neural spines, lack of pneumatic foramina on the lateral face of the centra and the presence of well-developed postexapophyses. Chaoyangopterids are, so far, represented with confidence only in Chinese deposits; the only record outside the Jehol Group is the Crato Formation form Lacusovagus magnificens , a partial skull whose assignment to the Chaoyangopteridae has been disputed. Given this controversy, we review the phylogenetic position of Lacusovagus , and discuss the nesting of our new specimen among theChaoyangopteridae, providing some comments concerning the composition of the group. We conclude that our new specimen provides further support for the presence of chaoyangopterids in the Early Cretaceous of Brazil. Supplementary material: Phylogenetic analysis data are available at: https://doi.org/10.6084/m9.figshare.c.3873391
FEEDING TRACES ON A PTERANODON (REPTILIA: PTEROSAURIA) BONE FROM THE LATE CRETACEOUS (CAMPANIAN) MOOREVILLE CHALK IN ALABAMA, USA
An istiodactylid pterosaur from the Upper Cretaceous Nanaimo Group, Hornby Island, British Columbia, Canada
New pterosaur tracks (Pteraichnidae) from the Late Cretaceous Uhangri Formation, southwestern Korea
A giant dromaeosaurid theropod from the Upper Cretaceous (Turonian) Bissekty Formation of Uzbekistan and the status of Ulughbegsaurus uzbekistanensis
Abstract: Six specimens accessioned to the Bavarian State Collection for Palaeontology and Geology in Munich, Germany, in 1966 are identified as coming from a gigantic pterodactyloid pterosaur. The previously undescribed material was obtained in 1955 by Jean Otto Haas and compares favourably in size with the type specimen of the Late Cretaceous (Maastrichtian) azhdarchid pterosaur Arambourgiania philadelphiae ( Arambourg 1959 ) from the same locality/region. The material represents fragments of two cervical vertebrae, a neural arch, a left femur, a ?radius, and a metacarpal IV and bones of problematic identity, and does not duplicate the type material of Arambourgiania . The timing of its collection and its locality of Ruseifa, Jordan suggest it might pertain to the same individual as the holotype.
Giant azhdarchid pterosaurs from the terminal Cretaceous of Transylvania (western Romania)
Abstract Pterosaur remains from the Late Cretaceous of the Haţeg Basin of western Romania were reported by Nopcsa as early as 1899. Recent discoveries from the Late Maastrichtian Densuş-Ciula Formation include the giant azhdarchid Hatzegopteryx thambema, the holotype of which, consisting of skull elements and a humerus from the Vălioara locality, is described in detail. A very large femur from the same formation at Tuştea is also described. The systematic position of Hatzegopteryx is discussed. The wing span of H. thambema is estimated to be close to that of Quetzalcoatlus northropi (>12 m), but its skull is especially robust and may have been remarkably long (> 2.5 m). The skull bones of H. thambema consist of a very thin outer cortex enclosing an inner meshwork of extremely thin trabeculae surrounding very numerous small alveoli, an unusual structure reminiscent of expanded polystyrene. This peculiar structure, combining strength with lightness, can probably be considered as an adaptation to flight in a very large animal, through reduction of skull weight.