... on the first distal articulated podomere of the frontal appendage ( Cong et al . 2018 ), and Innovatiocaris bears no such endite. Anomalocarididae, which is represented by its type species Anomalocaris canadensis , has endites of its frontal appendages with auxiliary spines on both anterior and posterior...

... in shaft # pd in distal articulated region tm between pd # rows en # of longest en en in d.a.r. alternate long/short as paired as increase in size from base to tip # large as small as between large as Amplectobeluidae 3 12/13 Yes 2 1 Yes Yes No 0/2 No Anomalocarididae 1–3 13...

... of Guanshan oral cone Large plates Small plates Furrowed folds Scale-like nodes Total steps A: Anomalocarididae + Amplectobeluidae 4 (1) 28 (1) Present or absent (1) Present or absent (1) 4 B: Hurdiidae 3 or 4 (1) 28 or more (1) Present (1) Present (1) 4 C: Basal lineage 4 (1) 28...

... Appendage KUMIP 204781a/b 5.3, 5.4 Wheeler Formation Cambrian, Drumian Peytoia nathorsti Hurdia Conway Morris and Robison, 1988 ; Daley et al., 2013a hurdiid Appendage KUMIP 312405a/b 3 Spence Shale Cambrian, Stage 5 Anomalocarididae gen. et sp. indet. Briggs et al., 2008 Hurdia...

...., 2021 ). It is incomplete proximally but likely comprises the distalmost podomere of the shaft, which bears a large hypertrophied endite, and an articulated region of 13 podomeres. It shows greatest similarity to Anomalocarididae and Amplectobeluidae, both of which include genera with paired endites...

... not correspond precisely to any known radiodont, but the appendage exhibits characters of both Anomalocarididae and Amplectobeluidae (see Cong et al., 2018 ; Pates and Daley, 2018 ; Guo et al., 2019 ). Two other animals are also new to the Parker Formation. One ( Fig. 2G ) is strongly reminiscent...

... canadensis , the type species of the nominate genus of Anomalocarididae, has not been comprehensively redescribed. Recent research has revealed that the anomalocaridids have higher taxonomic and morphological diversity than has been identified in the past, particularly in the Burgess Shale ( Daley and Budd...

...Figure 2 —Anomalocarididae gen. et sp. indet., Wheeler Formation, near Swasey Spring, House Range, Utah. 1 , KUMIP 314078, mouth apparatus, scale bar equals 1 cm. 2 , KUMIP 314086 (part), anterior appendage, scale bar equals 1 cm ...

... Amplectobeluidae, distant from the type species in Anomalocarididae, indicating they probably belong to a distinct genus. They are similar to the type species in their lack of posteriorly oriented auxiliary spines but easily differentiated by the outward kink separating the shaft and distal articulated regions...

.... 2014 ; VanRoy et al. 2015 ; Moysiuk and Caron 2019 ). Much of the morphological divergence between the four major radiodont morphogroups—Anomalocarididae, Amplectobeluidae, Tamisiocarididae, and especially Hurdiidae—is concentrated in the appendages, illustrated by their broad dispersal...

..., and are referred to the Paneuarthropoda following the convention described in Budd (2001) . Order Radiodonta Collins, 1996 Family Anomalocarididae Raymond, 1935 Several detailed studies of anomalocaridid morphology exist based primarily on material from the Burgess Shale and the Chengjiang...