Abstract This paper provides analysis of the published materials on the occurrences of the Dacian and Cimmerian molluscs in the Danube River valley as well as the results of Pliocene sediments study based on core material of the boreholes drilled at the Black Sea Shelf east of the Danube River Delta. In the early Pontian time, the Dacian Basin was a large sub-basin of Paratethys which, due to an abrupt drop in sea level, separated into the Euxinian, Dacian and Caspian basins. At the end of the Bosphorus time, the discharge of the Dacian Basin waters into the Euxinian Basin formed a wide valley from the Galati-Reni region to the east through the Galati gateway. During the Cimmerian transgression, a vast bay existed on the site of the modern Danube Delta, from which mutual migrations of the Dacian and Cimmerian molluscs took place along the runoff valley. The cessation of runoff occurred during the regressive phase of the Late Cimmerian. The rhythmically bedded thick strata originated during the existence of the runoff valley. These strata were identified as the Pridanubian Formation (Suite). The cryptogenic form of Tulotoma Tulotoma ( =Viviparus ) ovidii nasonis (Bogachev) is characteristic of the lower and middle parts of the suite. The presence of the Dacian and Cimmerian molluscs in this suite became the basis for the correlation of sediments of the Dacian and Cimmerian regional stages. The Duabian molluscs were registered in the Cimmerian deposits of the Transcaucasus (the Duabian layers), Priazovye and the Kerch–Taman region. The migration of these molluscs took place during the regressive phases due to the circular current in the Euxinian Basin similar to the one existing in the Black Sea today. The Pliocene formation contains marine and continental deposits of the Lower and Upper Pliocene, which are represented by the Pridanubian Formation (Lower and Upper), Cimmerian deposits (non-subdivided Lower and Middle Cimmerian), Lower Kujalnician deposits, Upper Poration deposits, complex of red-coloured palaeosols (the Upper Miocene–Lower Pliocene non-subdivided). The formation of the Pliocene sediments on the Black Sea Shelf, east of the Danube Delta, was controlled by the inter-basin connectivity of the Eastern Paratethys.