Abstract We present an updated look at Carboniferous brachiopod biozonation from most of the world framed into a revised Carboniferous palaeogeography, based on a selection of the literature published on Carboniferous brachiopods since the nineteenth century. The biostratigraphic significance of the most important brachiopod taxa is synthesized in seven geographical correlations. The Mississippian is characterized by rich brachiopod faunas, with widespread taxa with a good potential for global correlation, such as Rugosochonetes , Delepinea , Buxtonia , Antiquatonia , Spinocarinifera , Marginatia , Fluctuaria , Ovatia , Rhipidomella , Lamellosathyris , Unispirifer , Tylothyris and Syringothyris . From the mid-Visean to the late Serpukhovian, taxa of gigantoproductidines are biostratigraphically significant, and occur everywhere except South America and Australia, which remain as distinct faunal successions for most of the period. A major turnover occurs at the beginning of the Pennsylvanian, characterized by a higher degree of provincialism. Pennsylvanian brachiopod faunas are diverse in China, Russia and North America, but otherwise they are less developed and are characterized mostly by endemic taxa, hampering long-distance correlation. An exception is the rapid diversification of taxa of the Choristitinae, which were widespread from the Bashkirian to the Moscovian, allowing long-distance correlation.

Abstract: The Permian timescale has developed over about two centuries of research to the current chronostratigraphic scale advocated by the Subcommission on Permian Stratigraphy of three series and nine stages: Cisuralian (lower Permian) – Asselian, Sakmarian, Artinskian, Kungurian; Guadalupian (middle Permian) – Roadian, Wordian, Capitanian; and Lopingian (upper Permian) – Wuchiapingian and Changhsingian. The boundaries of the Permian System are defined by global stratotype sections and points (GSSPs) and the numerical ages of those boundaries appear to be determined with a precision better than 1‰. Nevertheless, much work remains to be done to refine the Permian timescale. Precise numerical age control within the Permian is very uneven and a global polarity timescale for the Permian is far from established. Chronostratigraphic definitions of three of the nine Permian stages remain unfinished and various issues of marine biostratigraphy are still unresolved. In the non-marine Permian realm, much progress has been made in correlation, especially using palynomorphs, megafossil plants, conchostracans and both the footprints and bones of tetrapods (amphibians and reptiles), but many problems of correlation remain, especially the cross-correlation of non-marine and marine chronologies. The further development of a Permian chronostratigraphic scale faces various problems, including those of stability and priority of nomenclature and concepts, disagreements over changing taxonomy, ammonoid v. fusulinid v. conodont biostratigraphy, differences in the perceived significance of biotic events for chronostratigraphic classification and correlation problems between provinces. Future research on the Permian timescale should focus on GSSP selection for the remaining undefined stage bases, the definition and characterization of substages, and further development and integration of the Permian chronostratigraphic scale with radioisotopic, magnetostratigraphic and chemostratigraphic tools for calibration and correlation.

Abstract: In 1841, Murchison coined the term Permian for strata in the Russian Urals. Recognition of the Permian outside of Russia and central Europe soon followed, but it took about a century for the Permian to be accepted globally as a distinct geological system. The work of the Subcommission on Permian Stratigraphy began in the 1970s and resulted in current recognition of nine Permian stages in three series: the Cisuralian (lower Permian) – Asselian, Sakmarian, Artinskian and Kungurian; the Guadalupian (middle Permian) – Roadian, Wordian and Capitanian; and the Lopingian (upper Permian) – Wuchiapingian and Changhsingian. The 1990s saw the rise of Permian conodont biostratigraphy, so that all Permian Global Stratigraphic Sections and Points (GSSPs) use conodont evolutionary events as the primary signal for correlation. Issues in the development of a Permian chronostratigraphic scale include those of stability and priority of nomenclature and concepts, disagreements over changing taxonomy, ammonoid v. fusulinid v. conodont biostratigraphy, differences in the perceived significance of biotic events for chronostratigraphic classification, and correlation problems between provinces. Further development of the Permian chronostratigraphic scale should focus on GSSP selection for the remaining, undefined stage bases, definition and characterization of substages, and further integration of the Permian chronostratigraphic scale with radioisotopic, magnetostratigraphic and chemostratigraphic tools for calibration and correlation.

Abstract: Establishing a Permian brachiopod biochronological scheme for global correlation is difficult because of strong provincialism during the Permian. In this paper, a brief overview of brachiopod successions in five major palaeobiogeographical realms/zones is provided. For Gondwanaland and peri-Gondwanan regions including Cimmerian blocks, Bandoproductus and Punctocyrtella (or Cyrtella ) are characteristic of the lower Cisuralian, as is Cimmeriella for the middle Cisuralian. As the Cimmerian blocks continued drifting north during the late Kungurian, accompanied by climate amelioration, contemporaneous brachiopods inhabiting these blocks showed a distinct shift from cold-water to mixed or warm-water affinities. However, coeval brachiopods in the Northern Transitional Zone (NTZ) are characterized by warm-water faunas and are associated with fusulinids in the lower Cisuralian. The Guadalupian brachiopods of the NTZ were clearly mixed between the Boreal and palaeoequatorial affinities. The end-Guadalupian is marked by the disappearance of a few characteristic genera, such as Vediproductus , Neoplicatifera and Urushtenoidea , in the Palaeotethyan region. The onset of the end-Permian mass extinction in the latest Changhsingian is clearly exhibited by the occurrence of the dwarfed and thin-shelled brachiopods commonly containing Paracrurithyris .