ABSTRACT Hotspots represent the ephemeral introduction of nutrients into an environment, and occur in both the modern and geologic past. The annual deposition of deciduous leaves in temperate forests, tree falls, animal excrement, and vertebrate carcass deposition all result in the pulsed introduction of nutrients to an ecosystem. Hotspots are critical for providing limiting nutrients, including nitrogen and carbon, to be incorporated into soil microbial biomass and plant biomass. For vertebrate ­carcasses, following the release of labile compounds from soft tissues, bones are often left behind, and provide a more recalcitrant reservoir of organic carbon and nitrogen, phosphorus, calcium, and, in some environments, water, for micro- and macro-fauna. Taphonomy—the physical, chemical, and biological processes following plant or animal death—studied in modern systems can be used to interpret hotspot processes operating in the past. East Tennessee is a region where studies of modern and fossil vertebrate hotspots have provided new insights into taphonomy. This guide describes two hotspot localities in east Tennessee—the Miocene-aged Gray Fossil Site in Gray, Tennessee, and the Anthropology Research Facility (“the Body Farm”) at the University of Tennessee, Knoxville, a human decomposition experimental site. The goal of this interdisciplinary field guide is to provide a view of nutrient hotspots from their formation in the modern to their preservation over geologic time.

Ceratopsid dinosaurs are notable for their common occurrences in bonebeds; however, until recently, these have not been encountered for the chasmosaurine Triceratops . The aim of this investigation is to describe the taphonomy of Quittin' Time (Museum of the Rockies locality HC-430), a Triceratops bonebed in the Hell Creek Formation, Garfield County, Montana. Using site taphonomic descriptions with an evaluation of ontogeny, inferences regarding the paleobiology of this extinct taxon are possible. The locality is associated with abundant organic material, including woody debris, large seeds, and other fragments in isolated silty lenses, all incorporated within a siltstone matrix, indicating preservation within a floodplain environment. Based on the repetition and ontogenetic stages of cranial elements, the minimum number of individuals (MNI) is three. Evidence from the location and taphonomic condition of the bones preserved in close proximity within the same siltstone unit suggests that the individuals—one young adult, one subadult, and a juvenile—likely accumulated during distinct flooding events within a narrow region of the floodplain as a result of “bloat-and-float” transport. The relatively small scale of the bonebed, both in terms of total area and number of individuals, implies that future work on Triceratops sites requires careful scrutiny of cranial elements examined within an ontogenetic framework because they are potentially critical to establishing MNI. Preservation of multiple individuals within the same unit does not necessarily provide evidence of gregarious behavior in Triceratops but rather may be a reflection of site taphonomic history and accumulation processes.